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conodont

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Dictionary: co·no·dont   ('nə-dŏnt', kŏn'ə-) pronunciation
n.
  1. A member of an extinct group of small primitive fishlike chordates, preserved primarily in the form of their conelike teeth.
  2. A fossil tooth of this chordate. Conodonts are the most widespread Paleozoic microfossils and are important for biostratigraphic indexing.

[Greek kōnos, cone + -ODONT.]


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Sci-Tech Encyclopedia: Conodont
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A group of extinct marine animals that are often abundant in strata of Late Cambrian to Late Triassic age, a time span of about 300 million years. Only the mineralized elements, which are usually 0.2 to 2 mm (0.008 to 0.08 in.) in dimension (the largest known reach 14 mm or 0.6 in.), are normally preserved. They are routinely extracted as isolated discrete specimens by chemical degradation of the rock in which they occur. In the earliest euconodonts (“true” conodonts, as opposed to the more primitive, and possibly unrelated, protoconodonts and paraconodonts), the elements comprise an upper crown and a basal body. The basal body occupies a cavity in the base of the crown, but is not present in the majority of post-Devonian species. In advanced conodonts the crown incorporates regular patches of opaque, finely crystalline, white matter.

For many years, conodont taxonomists treated individual element types as separate species. There are three major shape categories, coniform, ramiform, and pectiniform (see illustration). Coniform elements were dominant in the Cambrian to Early Ordovician and common until the Devonian. Ramiform (comblike) elements extend into elongate processes with various arrangements of denticles. Pentiniform elements include straight and arched blades, and may be expanded laterally to form a platform.

Conodont elements: (<i>a</i>, <i>b</i>) coniform elements, (<i>c</i>, <i>d</i>) ramiform elements, (<i>e</i>, <i>f</i>) pectiniform blade elements, (<i>g</i>, <i>h</i>) pectiniform platform elements, (<i>i</i>) bedding-plane <ailnk tname=assemblage.">
Conodont elements: (a, b) coniform elements, (c, d) ramiform elements, (e, f) pectiniform blade elements, (g, h) pectiniform platform elements, (i) bedding-plane assemblage.

In the absence of preserved soft parts, the nature of the affinities of conodonts was the subject of considerable speculation and debate. Since the first discovery of isolated elements in 1856, conodonts have been variously aligned with algae, higher plants, several wormlike phyla, mollusks, arthropods, lophophorates, chaetognaths, and chordates, or have been assigned to a separate phylum, Conodonta. It was not until 1983 that evidence of the soft parts was described by D. E. G. Briggs, E. N. K. Clarkson, and R. J. Aldridge, on the basis of the first of several specimens discovered in lower Carboniferous rocks near Edinburgh, Scotland. The evidence of the soft-part morphology indicates that the conodonts belong within the chordates; it is no longer possible to justify their separation as a phylum, Conodonta. See also Chordata.

Although the biological affinities of conodonts and the function of the elements were essentially unknown until recently, they have nonetheless been extensively studied because of their important geological applications. Most significant of these is the use of conodont elements in biostratigraphy. See also Stratigraphy.



Minute toothlike fossil composed of the mineral apatite (calcium phosphate); conodonts are among the most frequently encountered fossils in marine sedimentary rocks of Paleozoic age. They are the remains of animals that lived 543 – 248 million years ago that are believed to have been small marine invertebrates living in the open oceans and coastal waters throughout the tropical and temperate zones.

For more information on conodont, visit Britannica.com.

WordNet: conodont
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Note: click on a word meaning below to see its connections and related words.

The noun has 2 meanings:

Meaning #1: tiny fossil cone-shaped tooth of the most primitive vertebrate: the conodont

Meaning #2: small (2 in) extinct eellike fish with a finned tail and a notochord and having cone-shaped teeth containing cellular bone; late Cambrian to late Triassic; possible predecessor of the cyclostomes


Wikipedia: Conodont
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Conodonts
Fossil range: 495–199.6 Ma
Late Cambrian to Late Triassic

Reconstruction of a Conodont
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Class: Conodonta
Groups
  • Protoconodonta
  • Conodonta
    • Paraconodonta
    • Euconodonta
      • Panderodontida
      • Prioniodontida
        • Prioniodinina
        • Ozarkodinina
Conodont elements from the Deer Valley Member of the Mauch Chunk Formation

Conodonts are extinct chordates resembling eels, classified in the class Conodonta. For many years, they were known only from tooth-like microfossils now called conodont elements, found in isolation. The animals are also called Conodontophora (conodont bearers) to avoid ambiguity.

Contents

Description

The eleven known fossil imprints of conodont animals depict an eel-like creature with 15 or, more rarely, 19 elements forming a bilaterally symmetrical array in the head. This array constituted a feeding apparatus radically different from the jaws of modern animals. There are three forms of teeth, coniform cones, ramiform bars, and pectiniform platforms, which may have performed different roles.

The organisms range from a centimeter or so[verification needed] to the giant Promissum, 40 cm in length.[1] It is now widely agreed that conodonts had large eyes, fins with fin rays, chevron-shaped muscles and a notochord.

Ecology

The "teeth" of some conodonts have been interpreted as filter-feeding apparatuses, filtering out plankton from the water and passing it down the throat.[citation needed] Others have been interpreted as a "grasping and crushing array".[1]

The lateral position of the eyes makes a predatory role unlikely.[citation needed]

The preserved musculature hints that some conodonts (Promissum at least) were efficient cruisers but incapable of bursts of speed.[1]

Classification

The conodonts are currently classified in the phylum Chordata because their fins with fin rays, chevron-shaped muscles and notochord are characteristic of Chordata.[2]

They are considered by Milsom and Rigby to be vertebrates similar in appearance to modern hagfish and lampreys,[3] and phylogenetic analysis suggests that they are more derived than either of these groups.[4] This analysis, however, comes with one caveat: early forms of conodonts, the protoconodonts, appear to form a distinct clade from the later paraconodonts and euconodonts. It appears likely that the protoconodonts represent a stem group to the phylum containing chaetognath worms, indicating that they are not close relatives of true conodonts.[5]

Conodont teeth fossils

For many years, conodonts were known only from enigmatic tooth-like microfossils, which occur commonly but not always in isolation, and were not associated with any other fossil. These phosphatic microfossils are now termed "conodont elements" to avoid confusion. This confusion is most apparent for the non-specialist in the book "Your Inner Fish", by Neil Shubin, who describes the origin of teeth in chapter 4. In this chapter, the author attaches the name "conodont" to both the "conodont bearer" (the animal) and the "conodont elements" (the teeth), and the reader may have a hard time to make sense of the concept of "animals living in the mouths of animals".

They are widely used in biostratigraphy.

Conodont elements are also used as paleothermometers, a proxy for thermal alteration in the host rock. This is because under higher temperatures the phosphate undergoes predictable and permanent color changes, measured with the conodont alteration index. This has made them useful for petroleum exploration where they are known, in rocks dating from the Cambrian to the Late Triassic.

It was not until early 1980s that the conodont teeth were found in association with fossils of the host organism, in a konservat lagerstätte.[6] This is because most of the conodont animal was soft-bodied, thus everything but the teeth were not suited for preservation under normal circumstances.

Further reading

  • Aldridge, R. J., Briggs, D. E. G., Smith, M. P., Clarkson, E. N. K. & Clark, N. D. L. (1993), "The anatomy of conodonts". Philosophical Transactions of the Royal Society of London, Series B, 340, 405-421.
  • Aldridge, R. J. & Purnell, M. A. (1996). "The conodont controversies". Trends in Ecology and Evolution, 11, 463-468.
  • Donoghue, P. C. J., Forey, P. L. and Aldridge, R. J. (2000), "Conodont affinity and chordate phylogeny". Biological Reviews, 75, 191-251.
  • Janvier, P (1997). "Euconodonta". The tree of life web project, http://tolweb.org. http://tolweb.org/tree?group=Euconodonta&contgroup=Vertebrate. Retrieved 2007-09-05. 
  • Sweet, Walter. The Conodonta.
  • Sweet, W. C. and Donoghue, P. C. J. (2001), "Conodonts: past, present and future", Journal of Paleontology, 75, 1174-1184.

External links

References

  1. ^ a b c Gabbott, S.E.; R. J. Aldridge, J. N. Theron (1995). "A giant conodont with preserved muscle tissue from the Upper Ordovician of South Africa". Nature 374: 800–803. doi:10.1038/374800a0. 
  2. ^ Briggs, D. (May 1992). "Conodonts: a major extinct group added to the vertebrates". Science 256 (5061): 1285–1286. doi:10.1126/science.1598571. PMID 1598571. 
  3. ^ Milsom, Clare; Rigby, Sue (2004). "Vertebrates". Fossils at a Glance. Victoria, Australia: Blackwell Publishing. p. 88. ISBN 0632060476. 
  4. ^ Donoghue, P.C.J.; Forey, P.L.; Aldridge, R.J. (2000). "Conodont affinity and chordate phylogeny". Biological Reviews 75 (02): 191–251. doi:10.1017/S0006323199005472. http://journals.cambridge.org/abstract_S0006323199005472. Retrieved 2008-04-07. 
  5. ^ Szaniawski, H. (2002). "New evidence for the protoconodont origin of chaetognaths". Acta Palaeontologica Polonica 47 (3): 405. doi:10.1126/science.1137187. PMID 17332408. http://app.pan.pl/archive/published/app47/app47-405.pdf. 
  6. ^ Briggs, D.E.G.; E.N.K. Clarkson, R.J. Aldridge (1983). "The conodont animal". Lethaia 16: 1–14. doi:10.1111/j.1502-3931.1983.tb01139.x. 

 
 
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Jawless vertebrates
Hindeodus parvus
Promissum

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