Trees or shrubs in the genus Acer are commonly called Maples. They are variously classified in a family of their own, the
Aceraceae, or (together with the Hippocastanaceae)
included in the family Sapindaceae. Modern classifications, including the Angiosperm Phylogeny Group classification, favour inclusion in Sapindaceae. There are
approximately 125 species, most of which are native to Asia, but
several species also occur in Europe, northern Africa, and
North America.
The word Acer is derived from a Latin word meaning "sharp" (referring to the
characteristic points on the leaves) and was first applied to the genus by the French
botanist Joseph Pitton de Tournefort in 1700. The type species of the genus is
Acer pseudoplatanus (Sycamore Maple).[1]
Morphology
Maples are mostly trees growing to 10-40 metres (30-130 ft) in height. Others are
shrubs less than 10 metres tall with a number of small trunks originating at ground level. Most species are deciduous, but a
few in southern Asia and the Mediterranean region are evergreen. Most are shade-tolerant when young, and are often late-successional in ecology; many of the smaller
species are usually understory trees growing under the canopies of other larger trees, while the larger species eventually become
dominant canopy trees. Maple root systems are typically dense and fibrous. A few species, notably
Acer cappadocicum, frequently produce root
sprouts, which can develop into clonal colonies.[1]
Acer circinatum (Vine Maple) leaves showing the palmate veining typical of most
species
Maples are distinguished by opposite leaf arrangement. The leaves in most species are
palmate veined and lobed, with 3-9 (rarely to 13) veins each leading to a lobe, one of which
is central or apical. A small number of species differ in having palmate compound, pinnate
compound, pinnate veined or unlobed leaves. Several species, including Acer griseum
(Paperbark Maple), Acer mandshuricum (Manchurian Maple), Acer maximowiczianum (Nikko Maple), and Acer
triflorum (Three-flowered Maple), have trifoliate leaves. One species, Acer
negundo (Manitoba Maple), has pinnately compound leaves that may be simply trifoliate or may have five, seven, or
rarely nine leaflets. A few, such as Acer laevigatum and Acer carpinifolium (Hornbeam Maple), have pinnately-veined simple leaves.
The flowers are regular, pentamerous, and borne in
racemes, corymbs, or umbels.
They have four or five sepals, four or five petals about 1–6 mm
long (absent in some species), four to ten stamens about 6-10 mm long, and two pistils or a pistil with two styles. The ovary is superior and has two
carpels, whose wings elongate the flowers, making it easy to tell which flowers are female.
Maples flower in late winter or early spring, in most
species with or just after the leaves appear, but in some before them.[2]
Maple flowers are green, yellow, orange or red. Though individually small, the effect of an entire tree in flower can be
striking in several species. Some maples are an early spring source of pollen and
nectar for bees.
The distinctive fruit are called samaras or "maple
keys". These seeds occur in distinctive pairs each containing one seed enclosed in a "nutlet"
attached to a flattened wing of fibrous, papery tissue. They are shaped to spin as they fall and to carry the seeds a
considerable distance on the wind. Seed maturation is usually in a few weeks to six months after flowering, with seed dispersal
shortly after maturity. Most species require stratification in order to
germinate, and some seeds can remain dormant in the soil for several years before
germinating.[1]
The genus is subdivided by its morphology into a number of sections and subsections.[3]
Pests and diseases
Rhytisma acerinum fungus on
Acer pseudoplatanus leaf
The leaves are used as a food plant for the larvae of a number of Lepidoptera species (see List of Lepidoptera
which feed on Maples). Aphids are also very common sap-feeders on maples. In horticultural
applications a dimethoate spray will solve this.
Maples are affected by a number of fungal diseases. Several are susceptible to Verticillium
wilt caused by Verticillium species, which can cause significant local mortality.
Sooty bark disease, caused by Cryptostroma species, can kill trees which are under stress
due to drought. Death of maples can also be caused more rarely by Phytophthora root rot and Ganoderma root decay. Maple leaves
in late summer and autumn are commonly disfigured by "tar spot" caused by Rhytisma species
and mildew caused by Uncinula species, though these
diseases do not usually have an adverse effect on the trees' long-term health.[4]
Uses
Horticulture
Acer palmatum (Japanese Maple) has over 1,000
cultivars. This cultivar is
A. palmatum 'Sango kaku', sometimes called "coralbark maple".
Maples are planted as ornamental trees by homeowners, businesses and municipalities.
Acer platanoides (Norway Maple) is especially popular as it is fast-growing and
extremely cold-resistant, though is also an invasive species in some regions. Other
maples, especially smaller or more unusual species, are popular as specimen trees.[1]
Cultivars
Numerous maple cultivars have been selected for particular characteristics and can be
propagated only by grafting. Acer palmatum (Japanese Maple) alone has over 1,000 cultivars, most selected in Japan, and many of
them no longer propagated or not in cultivation in the western world. Some delicate
cultivars are usually grown in pots and rarely reach heights of more than 50-100 cm.[1]
Bonsai
Maples are a popular choice for the art of bonsai. Acer palmatum, Acer buergerianum (Trident Maple), Acer ginnala (Amur
Maple), Acer campestre (Field Maple) and Montpellier Maple (A. monspessulanum) are popular choices and respond well to techniques that
encourage leaf reduction and ramification, but most species can be used.[1]
Collections
Maple collections, sometimes called aceretums, occupy space in many gardens and arboreta around the world including the "five great W's" in England:
Wakehurst Place Garden, Westonbirt
Arboretum, Windsor Great Park, Winkworth
Arboretum and Wisley Garden. In the United
States, the aceretum at the Harvard-owned Arnold Arboretum in Boston is
especially notable. In the number of species and cultivars, the Esveld Aceretum in
Boskoop, Netherlands is the largest in the world.[1]
Tourism
Many maples have bright autumn foliage, and many countries have leaf-watching
traditions. In Japan, the custom of viewing the changing colour of maples in the autumn is called
"momijigari". Nikko and Kyoto are particularly favoured destinations for this activity.
The particularly spectacular fall colours of the Acer rubrum (Red Maple) are a
major contributor to the seasonal landscape in southeastern Canada and in New England. Fall tourism is a boon to the
economy of this region, especially in Vermont, New
Hampshire and Western Massachusetts. In the American Pacific Northwest, it is the spectacular fall colours of Acer
circinatum (Vine Maple) that draw tourists and photographers.
Commercial uses
Maples are important as source of syrup and wood. Dried wood is often used for the
smoking of food. They are also cultivated as ornamental plants and have benefits for tourism and agriculture.
Maple syrup
- Further information: Maple Syrup
The Sugar Maple (Acer saccharum) is tapped for sap, which is then boiled to produce maple syrup or made into
maple sugar or maple candy. It takes about 40 gallons
of Sugar Maple sap to make a gallon of syrup. Syrup can be made from closely-related species as well, but their output is
inferior. Sugar maples typically have a lifespan of 300 years.
Timber
A bench made of highly-figured maple wood
Some of the larger maple species have valuable timber, particularly Sugar Maple in North
America, and Sycamore Maple in Europe. Sugar Maple wood, often known as "hard maple", is the wood of choice for bowling pins, bowling alley lanes, drums and butcher's blocks. Maple wood is also used for the production of wooden baseball bats, though less often than ash or hickory.
Some maple wood has a highly decorative wood grain, known as flame maple and
quilt maple. This condition occurs randomly in individual trees of several species, and
often cannot be detected until the wood has been sawn, though it is sometimes visible in the standing tree as a rippled pattern
in the bark.
Maple is considered a tonewood, or a wood that carries sound waves well, and is used in
numerous musical instruments such as guitars and
drums. It provides resonance and a lighter weight than many other woods used in necks such as
rosewood. Also the look of a maple neck is appealing to many guitar players. Maple is also used to make bassoons.
Agriculture
As they are a major source of pollen in early spring before many other plants have flowered,
maples are important to the survival of honeybees that play a commercially-important role
later in the spring and summer.
Symbolism
The flag of Canada depicts a stylized maple leaf
and is a prominent national symbol. In the United States, the maple has been adopted by
New York,[5]
Vermont,[6]
Wisconsin[7] and
West Virginia.[8] The Red Maple was adopted by Rhode Island [9]as their official state
tree.
References
Wikimedia Commons has media related to:
hsb:Klonnds-nl:Plataan (Acer)
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