What is the primary difference between the Coptic Orthodox Church and the Greek Orthodox Church?

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Now there is no difference after many theological dialogue it came to the conclusion Greeks misunderstood the copts definition when describing the nature of christ.

In summary when the copts said christ had "one nature" they did not mean his divinity or humanity alone (which is what the Greeks thought) but the unity of both natures which formed one nature called "the logos incarnate",the copts rejected the Greeks definiton because they thought it was similair to the nestorious heresy.

Coptic point of view -
The Lord Jesus Christ is God Himself, the Incarnate Logos.Who took to Himself a perfect manhood. His Divine nature is one with his human nature yet without mingling, confusion or alteration; a complete Hypostatic Union. Words are inadequate to describe this union. It was said, that without controversy, "Great is the mystery of godliness: God was manifested in the flesh, (1 Tim. 3:16).As this union is permanent, never divided nor separated, we say in the liturgy that His Godhead never departed from His manhood for a single moment nor even for a twinkle of an eye. The Divine nature (God the Word) was united with the human nature which He took of the Virgin Mary by the action of the Holy Spirit. The Holy Spirit purified and sanctified the Virgin's womb so that the Child to whom she gave birth would inherit nothing of the original sin; the flesh formed of her blood was united with the Only-Begotten Son. This unity took place from the first moment of the Holy Pregnancy in the Virgin's womb.
As a result of the unity of both natures-the Divine and the human-inside the Virgin's womb, one nature was formed out of both: "The One Nature of God the Incarnate Logos" as St. Cyril called it.
The Holy Church did not find an expression more reliable, deep and precise than that which was used by St. Cyril the Great, and which St. Athanasius the Apostolic used before him. Both of them were true leaders in the theological field worldwide When the church participated in the dialogue arranged by the Pro Oriente group in Vienna, Austria in September 1971 between the Roman Catholic Church and the ancient Oriental Orthodox Churches concerning the Nature of Christ, the point of discussion was St. Cyril's expression "One Nature of God the Incarnate Logos" (Mia Physis Tou Theou Logou Sesarkwmene).After the schism which took place in the year 451 AD, when the Coptic Orthodox Church rejected the motions of the Council of Chalcedon and its theological struggles, we were called "Monophysites" that is, those who believe in the "One Nature".Sharing our belief are the Syrians, the Armenians, the Ethiopians and the Indians; who were also called "NonChalcedonian" Orthodox Churches.On the other hand, the Chalcedonian Catholic and Greek Churches "The Roman Orthodox" believe in the two natures of Christ; the Protestant Churches also hold this belief. Consequently, these churches are known as "Diophysites" - believers in the two natures of Christ. The Roman - or Chalcedonian - Orthodox Churches include those of Constantinople, Greece, Cyprus, Russia, Romania, Hungary and Serbia as well as the Roman Orthodox Churches of Egypt, Syria, Lebanon, America and the St. Catherine Monastery in the Sinai desert.The term "Monophysites" used for the believers in the One Nature has been intentionally or unintentionally misinterpreted throughout certain periods of history. Consequently, the Coptic and the Syrian Churches in particular were cruelly persecuted because of their belief, especially during the period which started from the Council of Chalcedon held in 451 AD and continued to the conquest of the Arabs in Egypt and Syria (about 641 AD).
This misinterpretation continued along history as though we believed in one nature of Christ and denied the other nature. We wonder which of the two natures the Church of Alexandria denies?
Is it the Divine nature? Certainly not, for our Church was the most fervent defender against the Arian heresy in the Council of Nicea, held in the year 325 AD, as well as before and after that. Or is it The Lord's human nature that the Church of Alexandria denies? St. Athanasius of Alexandria resolved this entirely in the oldest and greatest book on this subject The Incarnation of the Word,
The expression "One Nature" does not indicate the Divine nature alone nor the human nature alone, but it indicates the unity of both natures into One Nature which is "The Nature of the Incarnate Logos".
The same applies when we speak about our human nature which comprises two united natures: the soul and the body. Thus, man's nature is not the soul alone nor the body alone, but their union in one nature called human nature.
St. Cyril the Great taught us not to talk about two natures after their unity.So we can say that the Divine nature united hypostatically with the human nature within the Virgin's womb, but after this unity we do not ever speak again about two natures of Christ. In fact, the expression "two natures" implies in itself division or separation, and although those who believe in "the two natures" admit unity, the tone of separation was obvious in the Council of Chalcedon - a matter which prompted us to reject the Council and caused the exile of St. Dioscorus of Alexandria.
Since the Holy Synods of both the Coptic Orthodox Church and the Greek Orthodox Patriarchate of Alexandria and all Africa have already accepted the outcome of the official dialogue on Christology between the Orthodox Church and the Oriental Orthodox Churches, including the two official agreements: the first on Christology signed in June 1989 in Egypt and the second also on Christology and on the lifting of anathemas and restoration of full communion signed in Geneva 1990, in which it is stated that "In the light of our agreed statement on Christology..., we have now clearly understood that both families have always loyally maintained the same authentic Orthodox Christological faith, and the unbroken continuity of Apostolic tradition". It was agreed to have mutual recognition of the sacrament of Baptism, based on what St Paul wrote, "One Lord, one faith, one baptism" (Eph 4:5)
But since up until now we are waiting for the responses of the Holy Synods of some other churches in both families, the restoration of full communion is not yet reached between the two sides of the bi-lateral dialogue. And due to the pastoral consequences and implications caused by mixed Christian marriages between the members of the two Patriarchates of Alexandria, having the majority of their people living in the same countries. Those marriages being difficult to perform in both Churches at the same time or in concelebration. The result is that many sensitivities are created between the two families of the partners of such marriage. Those sensitivities which can extend even after the marriage and may affect the relation between the two communities of churches.
For those mentioned reasons, the Holy Synods of both Patriarchates have agreed to accept the sacrament of marriage which is conducted in either Church with the condition that it is conducted for two partners not belonging to the same Patriarchate of the other Church from their origin. Both the Bride and the Groom should carry a valid certificate from his/her own Patriarchate that he/she has a permit of marriage and indicating the details of his/her marriage status up to date.
Each of the two Patriarchates shall also accept to perform all of its other sacraments to that new family of Mixed Christian Marriage.
It is agreed that the Patriarchate which shall perform the marriage shall be responsible for any marriage problems that may happen concerning this certain marriage, taking into consideration the unified marriage laws signed by the heads of Churches in Egypt in the year 1999.
Each Patriarchate shall preserve its right not to give its sacraments to any persons whom she does not find fulfilling its canons according to the Apostolic Tradition.
Petros VII Pope and Patriarch of Alexandria and All Africa Shenouda III Pope of Alexandria and Patriarch of the See of St Mark .  The christology statement"We believe that our Lord, God and Saviour Jesus Christ, the Incarnate - Logos is perfect in His Divinity and perfect in His Humanity. He made His humanity One with His Divinity without Mixture, nor Mingling, nor Confusion. His Divinity was not separated from His Humanity even for a moment or twinkling of an eye. At the same time, we anathematize the Doctrines of both Nestorius and Eutyches".
Signatures.
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