2011 in paleontology

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2011 in paleontology

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Paleontology, palaeontology or palæontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because mankind has encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred in the year 2011.

Contents

Lobopods

Name Novelty Status Authors Age Unit Location Notes Images
Diania cactiformis[2]

Valid

Lower Cambrian

Maotianshan shale

 China

A lobopodian.

Mureropodia apae

Valid

  • Gamez Vintaned
  • Linan
  • Zhuravlev

Early Cambrian

Murero

 Spain

A lobopodian.

Siberion lenaicus

Valid

  • Dzik

Early Cambrian

Sinsk Formation

 Russia

A lobopodian.

Arthropods

Arachnids

Name Novelty Status Authors Age Unit Location Notes Images

Nephila jurassica

sp nov

valid

Selden, Shih & Ren

Middle Jurassic

Jiulongshan Formation

 China

largest fossil spider described

Insects

Name Novelty Status Authors Age Unit Location Notes Images

Agulla protomaculata[3]

sp nov

valid

Engel

Lutetian

Green River Formation

 USA

Only snakefly from the Green River Formation

Baltimartyria rasnitsyni[4]

sp nov

valid

Mey

Early Eocene

Baltic amber

 Europe

the second species in Baltimartyria

Diochus electrus[5]

sp nov

valid

Chatzimanolis & Engel

Lutetian

Baltic amber

 Europe

first valid Staphylininae species from the fossil record

Dryinus rasnitsyni[6]

sp nov

valid

Olmi & Guglielmino

Burdigalian

Dominican amber

 Dominican Republic

second fossil Dryinus lamellatus group species

Leptolingia calonervis[7]

sp.

Valid

  • Shi Chaofan
  • Yang Qiang
  • Ren Dong

Middle Jurassic

Jiulongshan Formation

 China

Leptolingia imminuta[8]

sp.

Valid

  • Yushuang Liu
  • Chaofan Shi
  • Dong Ren

Middle Jurassic

Daohugou Beds

 China

the smallest species known in the family Grammolingiidae

Litholingia ptesa[7]

sp.

Valid

  • Shi Chaofan
  • Yang Qiang
  • Ren Dong

Middle Jurassic

Jiulongshan Formation

 China

Myanmymar

gen et sp nov

valid

Huber

Albian

Burmese amber

 Myanmar

oldest mymarid described

Ordralfabetix[9]

gen et sp.

valid

Szwedo

Ypresian

Oise amber

 France

Second fossil lophopid planthopper from Europe

Platyperla marquati[10]

Species

Valid

Late Triassic

Potrerillos Formation

A platyperlid stonefly, a species of Platyperla.

Plumalexius[11]

fam, gen et sp.

vaild

Brothers

Turonian

New Jersey amber

 USA

sister genus to Plumariidae

Renphasma[12]

Gen. et sp.

Valid

  • Nel
  • Delfosse

Early Cretaceous

Yixian Formation

 China

A stick insect

Titanomyrma lubei[13]

Gen. et sp.

valid

  • Archibald
  • Johnson
  • Mathewes
  • Greenwood

Early Eocene

Green River Formation USA
Messel shales, Germany
Eckfeld Maar, Germany

 USA
 Germany

new genus for the largest known ants

Mickoleitia longimanus[14]

ord., fam., gen. et sp.

  • Staniczek
  • Bechly
  • Godunko

Lower Cretaceous

Crato Formation

 Brazil

A strange stem group representative of mayflies that has been attributed to a new family Mickoleitiidae and a new fossil insect order Coxoplectoptera.

Fishes

Newly named jawless vertebrates

Name Novelty Status Authors Age Unit Location Notes Images

Althaspis tarloi[15]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Djurinaspis secunda[15]

sp.

Valid

  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Lechriaspis[16]

Gen. et sp.

Valid

  • Elliott
  • Petriello

Early Devonian

Water Canyon Formation

A heterostracan

Palanasaspis[15]

Gen. et sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Podolaspis danieli[15]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A heterostracan

Poraspis thomasi[16]

Species

Valid

  • Elliott
  • Petriello

Early Devonian

Lost Burro Formation

A heterostracan

Semipodolaspis[15]

Gen. et sp.

Valid

  • Voichyshyn

Early Devonian

Tyver Formation

A heterostracan

Shuyu[17]

Gen. et comb.

Valid

Late Telychian/Early Wenlock

Maoshan Formation

The basalmost known eugaleaspidiform galeaspid.

Zenaspis kasymyri[15]

sp.

Valid

  • Voichyshyn

Early Devonian

Dniester Formation

A osteostracan

Newly named acanthodian

Name Novelty Status Authors Age Unit Location Notes Images

Nerepisacanthus[18]

Gen. et sp.

Valid

  • Burrow

?Telychian/Ludlow

Cunningham Creek Formation

An acritolepid ischnacanthiform.

Newly named cartilaginous fishes

Name Novelty Status Authors Age Unit Location Notes Images

Dwardius sudindicus[19]

Species

Valid

Early Cenomanian

Karai Formation

An otodontid.

Fayolia sharovi[20]

Species

Valid

Middle/Late Triassic

Madygen Formation

Egg capsules of a probable xenacanthid.

Gladioserratus[19]

Gen. et comb.

Valid

  • Underwood
  • Goswami
  • et al.

Hauterivian to Cenomanian

A cow shark, a new genus for "Notidanus" aptiensis (Pictet, 1865).

Gladioserratus magnus[19]

Species

Valid

  • Underwood
  • Goswami
  • et al.

Early Cenomanian

Karai Formation

A cow shark, a second species of Gladioserratus.

Kawichthys[21]

Gen. et sp.

Valid

Late Virgilian

Douglas Group

A symmoriiform.

Lonchidion ferganensis[20]

Species

Valid

  • Fischer
  • Voigt
  • et al.

Middle/Late Triassic

Madygen Formation

A hybodontid shark.

Palaeoxyris alterna[20]

Species

Valid

  • Fischer
  • Voigt
  • et al.

Middle/Late Triassic

Madygen Formation

A hybodontid shark.

Reginaselache[22]

Gen. et sp.

Valid

  • Turner
  • Burrow

Middle Viséan

Ducabrook Formation

A xenacanthiform.

Tassiliodus[23]

Gen. et sp.

Valid

  • Derycke
  • Goujet

Emsian, Devonian

An euselachian.

Newly named bony fishes

Name Novelty Status Authors Age Unit Location Notes Images

Araripichthys weberi[24]

Species

Valid

  • Alvarado-Ortega
  • Brito

Albian

Tlayúa Formation

An elopocephalan, a species of Araripichthys.

Beishanichthys[25]

Gen. et sp.

Valid

  • Xu
  • Gao

Early Triassic

A scanilepiform.

Bourbonnella jocelynae[26]

Species

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

An aeduellid.

Caucasichthys[27]

Gen. et sp.

Valid

  • Bannikov
  • Carnevale
  • Parin

Bartonian

A perciform.

Dapedium stollorum[28]

Species

Valid

  • Thies
  • Hauff

Early Toarcian

A species of Dapedium.

Guntherichthys[26]

Gen. et sp.

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Laccognathus embryi[29]

Species

Valid

Frasnian

Fram Formation

A porolepiform sarcopterygian.

Lepidotes pankowskii[30]

Species

Valid

  • Forey
  • López-Arbarello
  • MacLeod

Cenomanian

Kem Kem Beds

A species of Lepidotes.

Leufuichthys[31]

Gen. et sp.

Valid

  • Gallo
  • Calvo
  • Kellner

Turonian/Coniacian

Portezuelo Formation

A clupeomorph.

Mimipiscis[32]

Nom. nov.

Valid

  • Choo

Upper Devonian

Gogo Formation

A new generic name for Mimia Gardiner & Bartram (1977). The type species is Mimipiscis toombsi (Gardiner & Bartram 1977).

Mimipiscis bartrami[32]

Species

Valid

  • Choo

Upper Devonian

Gogo Formation

A second species of Mimipiscis.

Pietschellus[33]

Gen. et sp.

Valid

  • Bannikov
  • Carnevale

Ypresian

Monte Bolca

A percomorph.

Scheenstia[34]

Gen. et sp.

Valid

  • López-Arbarello
  • Sferco

Late Jurassic

A semionotiform.

Sinosaurichthys[35]

Gen. et 3 sp.

Valid

Anisian

Guanling Formation

A saurichthyid.

Sorbinichthys africanus[36]

Species

Valid

  • Murray
  • Wilson

Cenomanian or Turonian

Akrabou Formation

A species of Sorbinichthys.

Spinofacia[26]

Gen. et sp.

Valid

  • Mickle

Carboniferous

Manning Canyon Shale Formation

A ’palaeoniscoid’.

Amphibians

Newly named lepospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Kirktonecta[37]

Gen. et sp.

Valid

  • Clack

Viséan

East Kirkton Quarry

A microsaur.

Tambaroter[38]

Gen. et sp.

Valid

Early Permian

Tambach Formation

An ostodolepid.

Newly named temnospondyls

Name Novelty Status Authors Age Unit Location Notes Images

Acheloma dunni[39]

Species

Valid

  • Polley
  • Reisz

Early Permian

A trematopid.

Calmasuchus[40]

Gen. et sp.

Valid

  • Fortuny
  • Galobart
  • De Santisteban

early-middle Anisian

Catalonian basin

A capitosaurian.

Rotaryus[41]

Gen. et sp.

Valid

Early Permian

Tambach Formation

A trematopid.

Rubeostratilia[42]

Gen. et sp.

Valid

  • Bourget
  • Anderson

Early Permian

An amphibamid.

Newly named lissamphibians

Name Novelty Status Authors Age Unit Location Notes Images

Urupia[43]

Gen. et sp.

Valid

  • Skutschas
  • Krasnolutskii

Bathonian

Itat Formation

A basal stem salamander.

Anapsids

Newly named basal anapsids

Name Novelty Status Authors Age Unit Location Notes Images

Gansurhinus[44]

Gen. et sp.

Valid

Middle Permian

A moradisaurine captorinid.

Newly named turtles

Name Novelty Status Authors Age Unit Location Notes Images

Albertwoodemys[45]

Gen. et sp.

Valid

Early Oligocene

Jebel Qatrani Formation

A podocnemidid.

Brontochelys[45]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

A podocnemidid, a new genus for "Shweboemys" gaffneyi.

Cordichelys[45]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Eocene

Qasr el-Sagha beds

A podocnemidid, a new genus for "Podocnemis" antiqua.

Gilmoremys[46]

Gen. et comb.

Valid

  • Joyce
  • Lyson

Maastrichtian

Lance Formation
Hell Creek Formation

A softshell turtle, new genus for "Aspideretes" lancensis (Gilmore, 1928).

Hispaniachelys[47]

Gen. et sp.

Valid

Oxfordian

Lorente Formation

A paracryptodire or a basal testudine.

Hoplochelys clark[48]

Species

Valid

Maastrichtian

Hell Creek Formation

A kinosternoid, a species of Hoplochelys.

Lapparentemys[45]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Paleocene

Santa Lucía Formation

A podocnemidid, a new genus for "Roxochelys" vilavilensis.

Latentemys[45]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Uncertain, possibly Miocene.

Uncertain, possibly Moghara Formation.

A podocnemidid.

Lemurchelys[45]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Early Miocene

Moghara Formation

A podocnemidid.

Mogharemys[45]

Gen. et comb.

Valid

  • Gaffney
  • Meylan
  • et al.

Miocene

A podocnemidid, a new genus for "Sternothaerus" blanckenhorni.

Oliveremys[49]

Gen. et comb. nov.

Valid

  • Vitek

Eocene

Washakie Formation

A trionychid, a new genus for "Trionyx" uintaensis.

Patagoniaemys[50]

Gen. et sp.

Valid

  • Sterli
  • De la Fuente

Campanian - Maastrichtian

La Colonia Formation

A stem turtle.

Peiropemys[45]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Pricemys[45]

Gen. et sp.

Valid

  • Gaffney
  • Meylan
  • et al.

Late Cretaceous

Marília Formation

A podocnemidid.

Prochelidella cerrobarcinae[51]

Species

Valid

Aptian/Albian

Cerro Barcino Formation

The oldest known chelid, a species of Prochelidella.

Selenemys[52]

Gen. et sp.

Valid

  • Pérez-García
  • Ortega

Upper Kimmeridgian

Lusitanian Basin

A pleurosternid.

Sokatra[53]

Gen. et sp.

Valid

  • Gaffney
  • Krause

Maastrichtian

Maevarano Formation

A pleurodiran.

Trionyx ikoviensis[54]

Species

Valid

Middle Eocene

A species of Trionyx sensu lato.

Diapsids

Newly named basal diapsids

Name Novelty Status Authors Age Unit Location Notes Images

Orovenator[55]

Gen. et sp.

Valid

Sakmarian

The oldest and most basal neodiapsid.

Newly named ichthyosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Sveltonectes[56]

Gen. et sp.

Valid

Barremian

An ophthalmosaurid.

Lepidosauromorphs

Newly named saurosphargids

Name Novelty Status Authors Age Unit Location Notes Images

Sinosaurosphargis[57]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

An incertae sedis.

Newly named sauropterygians

Name Novelty Status Authors Age Unit Location Notes Images

Abyssosaurus[58]

Gen. et sp.

Valid

  • Berezin

Late Hauterivian

An aristonectid.

Diandongosaurus[59]

Gen. et sp.

Valid

  • Shang
  • Wu
  • Li

Middle Anisian

Guanling Formation

A relative of nothosauroids, pachypleurosaurs and Wumengosaurus.

Dianopachysaurus[60]

Gen. et sp.

Valid

Middle Anisian

Guanling Formation

A keichousaurid pachypleurosaur.

Hauffiosaurus tomistomimus[61]

Species

Valid

Early Toarcian

Whitby Limestone Formation

A second species of Hauffiosaurus.

Marmornectes[62]

Gen. et sp.

Valid

  • Ketchum
  • Benson

Callovian

Oxford Clay Formation

A pliosaurid.

Westphaliasaurus[63]

Gen. et sp.

Valid

  • Schwermann
  • Sander

Pliensbachian

Höxter district

A plesiosaurid plesiosaur.

Zarafasaura[64]

Gen. et sp.

Valid

Maastrichtian

An elasmosaurid plesiosaur.

Newly named lizards

Name Novelty Status Authors Age Unit Location Notes Images

Brasiliguana[65]

Gen. et sp.

Valid

  • Nava
  • Martinelli

Turonian/Santonian

Adamantina Formation

A iguanian lizard.

Cryptolacerta[66]

Gen. et sp.

Valid

Lutetian

Messel pit

A amphisbaenian lizard.

Latoplatecarpus[67]

Gen. et sp.

Valid

  • Konishi
  • Caldwell

Campanian

Pierre Shale Formation
Demopolis Chalk Formation

A plioplatecarpine mosasaur. Its type species is Latoplatecarpus willistoni; "Plioplatecarpus" nichollsae (Cuthbertson et al., 2007) is a second species of Latoplatecarpus.

Plesioplatecarpus[67]

Gen. et comb.

Valid

  • Konishi
  • Caldwell

Middle Coniacian to middle Santonian.

Niobrara Chalk
Eutaw Formation

A plioplatecarpine mosasaur, a new genus for "Clidastes" planifrons (Cope, 1874).

Newly named snakes

Name Novelty Status Authors Age Unit Location Notes Images

Madtsoia pisdurensis[68]

Species

Valid

  • Mohabey
  • Head
  • Wilson

Late Cretaceous

A madtsoiid snake.

Archosauromorphs

Newly named basal archosauromorphs

Name Novelty Status Authors Age Unit Location Notes Images

Archeopelta[69]

Gen. et sp.

Valid

  • Desojo
  • Ezcurra
  • Schultz

Late Ladinian/Early Carnian

Santa Maria Formation

A doswelliid

Augustaburiania[70]

Gen. et sp.

Valid

  • Sennikov

Early Triassic

Don River Basin

The earliest tanystropheid.

Kyrgyzsaurus[71]

Gen. et sp.

Valid

  • Alifanov
  • Kurochkin

Late Ladinian/Early Carnian

Madygen Formation

The earliest drepanosaurid.

Protanystropheus[70]

Gen. et comb.

Valid

  • Sennikov

Middle - Late Triassic

A tanystropheid, a new genus for "Tanystropheus" antiquus (von Huene, 1905).

Newly named crurotarsans

Name Novelty Status Authors Age Unit Location Notes Images

Acherontisuchus[72]

Gen. et sp.

Valid

  • Hastings
  • Bloch
  • Jaramillo

Middle/Late Paleocene

Cerrejón Formation

A dyrosaurid.

Anteophthalmosuchus[73]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid.

Arenysuchus[74]

Gen. et sp.

Valid

  • Puértolas
  • Canudo
  • Cruzado-Caballero

Maastrichtian

Tremp Formation

A stem crocodyloid.

Caipirasuchus[75]

Gen. et sp.

Valid

  • Iori
  • Carvalho

Turonian/Santonian

Adamantina Formation

A sphagesaurid.

Campinasuchus[76]

Gen. et sp.

Valid

Turonian/Santonian

Adamantina Formation

A baurusuchid.

Caryonosuchus[77]

Gen. et sp.

Valid

Campanian/Maastrichtian

Adamantina Formation

A sphagesaurid.

Decuriasuchus[78]

Gen. et sp.

Valid

  • França
  • Ferigolo
  • Langer

Ladinian

Santa Maria Formation

Skeletons indicate possible group behavior.

Fruitachampsa[79]

Gen. et sp.

Valid

  • Clark

Late Jurassic

Morrison Formation

A shartegosuchid.

Goniopholis kiplingi[80]

Species

Valid

Berriasian

Purbeck Limestone Group

A goniopholidid.

Goniopholis willetti[73]

Species

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A goniopholidid.

Labidiosuchus[81]

Gen. et sp.

Valid

Maastrichtian

Marília Formation

A notosuchian.

Leiokarinosuchus[73]

Gen. et sp.

Valid

  • Salisbury
  • Naish

Early Cretaceous

Wealden Group

A neosuchian.

Lorosuchus[82]

Gen. et sp.

Valid

  • Pol
  • Powell

Paleocene

Rio Loro Formation

A sebecid.

Neptunidraco[83]

Gen. et sp.

Valid

  • Cau
  • Fanti

late Bajocian/early Bathonian

Rosso Ammonitico Veronese Formation

The oldest known metriorhynchid.

Pepesuchus[84]

Gen. et sp.

Valid

Campanian/Maastrichtian

Presidente Prudente Formation

A peirosaurid.

Pietraroiasuchus[85]

Gen. et sp.

Valid

Early Albian

Pietraroia Plattenkalk

A hylaeochampsid.

Pissarrachampsa[86]

Gen. et sp.

Valid

  • Montefeltro
  • Larsson
  • Langer

Campanian/Maastrichtian

Vale do Rio do Peixe Formation

A baurusuchid. The type species is Pissarrachampsa sera.

Theriosuchus grandinaris[87]

Species

Valid

Berriasian/Barremian

Khua Formation

A fifth species of Theriosuchus.

Newly named birds

Name Novelty Status Authors Age Unit Location Notes Images

Bohaiornis[88]

Gen. et sp.

Valid

Barremian/Aptian

Yixian Formation

An enantiornithine.

Cyrilavis colburnorum[89]

Species

Valid

  • Ksepka
  • Clarke
  • Grande

Eocene

Green River Formation

A halcyornithid stem parrot, a species of Cyrilavis.

Gracilornis[90]

Gen. et sp.

Valid

  • Li
  • Hou

Aptian

Jiufotang Formation

An enantiornithine.

Hoazinavis[91]

Gen. et sp.

Valid

  • Mayr
  • Alvarenga
  • Mourer-Chauviré

Late Oligocene/Early Miocene

Tremembé Formation

An early hoatzin.

Itaboravis[92]

Gen. et sp.

Valid

  • Mayr
  • Alvarenga
  • Clarke

Early late Paleocene (Itaboraian).

A possible member of Cariamae. The type species is Itaboravis elaphrocnemoides.

Jamna[93]

Gen. et sp.

Valid

Oligocene

Carpathian flysch

A passerine.

Kaiika[94]

Gen. et sp.

Valid

  • Fordyce
  • Thomas

Early Eocene

Kauru Formation

A basal penguin.

Lavocatavis[95]

Gen. et sp.

Valid

  • Mourer-Chauviré
  • Tabuce
  • et al.[CAL 28]

Eocene

Glib Zegdou Formation

A phorusrhacoid.

Mystiornis[96]

Gen. et sp.

Valid

Barremian/Aptian

Shestakovo-1 locality

Nambashag[97]

Gen. et 2 sp.

Valid

  • Worthy

Late Oligocene/Early Miocene

Etadunna Formation
Namba Formation

A cormorant.

Parahongshanornis[98]

Gen. et sp.

Valid

  • Li
  • Wang
  • Hou

Early Cretaceous

Jiufotang Formation

A hongshanornithid.

Qiliania[99]

Gen. et sp.

Valid

Aptian

Xiagou Formation

An enantiornithine.

Newly named non-avian dinosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Acristavus[100]

Gen. et sp.

Valid

Campanian

Two Medicine Formation
Wahweap Formation

A brachylophosaurin hadrosaurine.

Ahshislepelta[101]

Gen. et sp.

Valid

  • Burns
  • Sullivan

Late Campanian

Kirtland Formation

An ankylosaurine ankylosaurid.

Albinykus[102]

Gen. et sp.

Valid

Late Cretaceous

An alvarezsaurid theropod.

Amtocephale[103]

Gen. et sp.

Valid

  • Watabe
  • Tsogtbaatar
  • Sullivan

Turonian-Santonian

Baynshire Formation

A pachycephalosaurid.

Angolatitan[104]

Gen. et sp.

Valid

Late Turonian

Itombe Formation

A somphospondylian sauropod.

Arcusaurus[105]

Gen. et sp.

Valid

  • Yates
  • Bonnan
  • Neveling

Lower Jurassic

Elliot Formation

Basal Sauropodomorph

Atacamatitan[106]

Gen. et sp.

Valid

Late Cretaceous

Tolar Formation

A titanosaur.

Brontomerus[107]

Gen. et sp.

Valid

  • Taylor
  • Wedel
  • Cifelli

Aptian/Albian

Cedar Mountain Formation

A camarasauromorph sauropod.

Daemonosaurus[108]

Gen. et sp.

Valid

Late Triassic

Chinle Formation

A primitive theropod.

Delapparentia[109]

Gen. et sp.

Valid

  • Ruiz-Omeñaca

Early Barremian

Camarillas Formation

An iguanodontoid.

Demandasaurus[110]

Gen. et sp.

Valid

  • Fernández-Baldor
  • Canudo
  • et al.[CAL 36]

Late Barremian-early Aptian

Castrillo la Reina Formation

A rebbachisaurid sauropod.

Drusilasaura[111]

Gen. et sp.

Valid

  • Navarrete
  • Casal
  • Martínez

Cenomanian/Turonian

Bajo Barreal Formation

A possible lognkosaurian titanosaur.

Eodromaeus[112]

Gen. et sp.

Valid

Middle Carnian

Ischigualasto Formation

A basal theropod.

Epichirostenotes[113]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Late Campanian

Horseshoe Canyon Formation

A caenagnathid.

Haya[114]

Gen. et sp.

Valid

Late Cretaceous

Javkhlant Formation

Basal ornithopod with many gastroliths.

Jaklapallisaurus[115]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation
Lower Dharmaram Formation

A plateosaurid.

Koreaceratops[116]

Gen. et sp.

Valid

  • Lee
  • Ryan
  • Kobayashi

Albian

"Tando beds"

A basal ceratopsian.

Koreanosaurus[117]

Gen. et sp.

Valid

?Santonian/Campanian

Seonso Conglomerate

A hypsilophodontid ornithopod.

Leonerasaurus[118]

Gen. et sp.

Valid

  • Pol
  • Garrido
  • Cerda

Pliensbachian/Toarcian

Las Leoneras Formation

An anchisaurian sauropodomorph.

Leyesaurus[119]

Gen. et sp.

Valid

Lower Jurassic

Quebrada del Barro Formation

A massospondylid sauropodomorph.

Linhenykus[120]

Gen. et sp.

Valid

Campanian

Wulansuhai Formation

An alvarezsaurid.

Linhevenator[121]

Gen. et sp.

Valid

Campanian

Bayan Mandahu Formation

An short-armed troodontid.

Manidens[122]

Gen. et sp.

Vald

  • Pol
  • Rauhut
  • Becerra

Aalenian/Early Bathonian

Cañadón Asfalto Formation

A heterodontosaurid.

Nambalia[115]

Gen. et. sp.

Valid

Late Norian-earliest Rhaetian

Upper Maleri Formation

A basal sauropodomorph.

Narambuenatitan[123]

Gen. et sp.

Valid

  • Filippi
  • García
  • Garrido

lower-middle Campanian

Anacleto Formation

A lithostrotian titanosaur.

Ojoraptorsaurus[113]

Gen. et sp.

Valid

  • Sullivan
  • Jasinski
  • Van Tomme

Lower Maastrichtian

Ojo Alamo Formation

A caenagnathid.

Omeisaurus jiaoi[124]

Species

Valid

Middle Jurassic

Xiashaximiao Formation

A species of Omeisaurus.

Osmakasaurus[125]

Gen. et comb.

Valid

  • McDonald

Valanginian

Lakota Formation

A new genus for "Camptosaurus" depressus (Gilmore, 1909).

Oxalaia[126]

Gen. et sp.

Valid

Cenomanian

Alcântara Formation

Huge spinosaur 12–14 m long.

Pampadromaeus[127]

Gen. et sp.

Valid

Carnian

Santa Maria Formation

A very basal sauropodomorph.

Pamparaptor[128]

Gen. et sp.

Valid

  • Porfiri
  • Calvo
  • Santos

Turonian/Coniacian

Portezuelo Formation

A deinonychosaur.

Petrobrasaurus[129]

Gen. et sp.

Valid

Santonian

Plottier Formation

A titanosaur.

Propanoplosaurus[130]

Gen. et sp.

Valid

  • Stanford
  • Weishampel
  • Deleon

Early Aptian

Patuxent Formation

A nodosaurid.

Qiupalong[131]

Gen. et sp.

Valid

Late Cretaceous

Qiupa Formation

An ornithomimid.

Ratchasimasaurus[132]

Gen. et sp.

Valid

  • Shibata
  • Jintasakul
  • Azuma

Aptian

Khok Kruat Formation

A non-hadrosaurid iguanodontian.

Sarahsaurus[133]

Gen. et sp.

Valid

Sinemurian/Pliensbachian

Kayenta Formation

A basal sauropodomorph

Siamodon[134]

Gen. et sp.

Valid

  • Buffetaut
  • Suteethorn

Aptian

Khok Kruat Formation

An advanced iguanodontian.

Spinops[135]

Gen. et sp.

Valid

Campanian

Oldman or Dinosaur Park Formation

A centrosaurine ceratopsian.

Stegoceras novomexicanum[136]

Species

Valid

  • Jasinski
  • Sullivan

Late Campanian

Fruitland Formation
Kirtland Formation

Second known species of Stegoceras.

Talos[137]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A troodontid theropod.

Tapuiasaurus[138]

Gen. et sp.

Valid

Aptian

Quiricó Formation

A titanosaur.

Teratophoneus[139]

Gen. et sp.

Valid

Late Campanian

Kaiparowits Formation

A tyrannosaurine tyrannosaurid.

Thescelosaurus assiniboiensis[140]

Species

Valid

  • Brown
  • Boyd
  • Russell

Late Maastrichtian

Frenchman Formation

Third known species of Thescelosaurus.

Titanoceratops[141]

Gen. et sp.

Valid

  • Longrich

Late Campanian

Fruitland or Kirtland Formation

A giant chasmosaurine ceratopsian from New Mexico.

Traukutitan[142]

Gen. et sp.

Valid

  • Juárez Valieri
  • Calvo

Santonian

Bajo de la Carpa Formation

A possible lognkosaurian titanosaur.

Uteodon[125]

Gen. et comb.

Valid

  • McDonald

Late Jurassic

Morrison Formation

A new genus for "Camptosaurus" aphanoecetes (Carpenter & Wilson, 2008).

Veterupristisaurus[143]

Gen. et sp.

Valid

  • Rauhut

Late Kimmeridgian/earliest Tithonian

Tendaguru Formation

A carcharodontosaurid.

Xiaotingia[144]

Gen. et sp.

Valid

early Late Jurassic

Tiaojishan Formation

An Archaeopteryx-like paravian.

Xuwulong[145]

Gen. et sp.

Valid

  • You
  • Li
  • Liu

Early Cretaceous

Xinminpu Group

A hadrosauriform.

Zhuchengtyrannus[146]

Gen. et sp.

Valid

Late Cretaceous

Wangshi Group

A tyrannosaurine tyrannosaurid.

Newly named pterosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Aurorazhdarcho[147]

Gen. et sp.

Valid

  • Frey
  • Meyer
  • Tischlinger

Tithonian

Solnhofen limestone

An azhdarchoid.

Aussiedraco[148]

Gen. et sp.

Valid

  • Kellner
  • Rodrigues
  • Costa

Albian

Toolebuc Formation

An ornithocheiroid.

Barbosania[149]

Gen. et sp.

Valid

  • Elgin
  • Frey

Cenomanian

Santana Formation

An ornithocheirid.

Darwinopterus robustodens[150]

Species

Valid

Middle Jurassic

Tiaojishan Formation

A darwinopterid.

Gwawinapterus[151]

Gen. et sp.

Valid

  • Arbour
  • Currie

Campanian

Northumberland Formation

An istiodactylid.

Jianchangopterus[152]

Gen. et sp.

Valid

  • Bo

Middle Jurassic

Tiaojishan Formation

A rhamphorhynchid.

Navajodactylus[153]

Gen. et sp.

Valid

  • Sullivan
  • Fowler

Late Campanian

Kirtland Formation
Dinosaur Park Formation

A possible azhdarchid.

Pterofiltrus[154]

Gen. et sp.

Valid

  • Jiang
  • Wang

Early Cretaceous

Yixian Formation

A ctenochasmatid.

Unwindia[155]

Gen. et sp.

Valid

  • Martill

Cenomanian

Santana Formation

A basal ctenochasmatoid.

Synapsids

Newly named non-mammalian synapsids

Name Novelty Status Authors Age Unit Location Notes Images

Annatherapsidus postum[156]

Sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Basilodon[157]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" woodwardi.

Bridetherium[158]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Candelariodon[159]

Gen. et sp.

Valid

Middle Triassic

Santa Maria Formation

A carnivorous cynodont.

Cryptovenator[160]

Gen. et sp.

Valid

Late Gzhelian

Remigiusberg Formation

A sphenacodontid pelycosaur. The type species is Cryptovenator hirschbergeri.

Euptychognathus[157]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" bathyrhynchus.

Euromycter[161]

Gen. et comb.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. A new genus for "Casea" rutena.

Gorochovetzia[156]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Keyseria[157]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" benjamini.

Muchia[156]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian.

Paceyodon[158]

Gen. et sp.

Valid

  • Clemens

Early Jurassic

A morganucodontan.

Peramodon[157]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" amalitzkii

Perplexisaurus lepusculus[156]

Species

Valid

  • Ivakhnenko

Middle Permian

A therocephalian.

Purlovia[156]

Gen. et sp.

Valid

  • Ivakhnenko

Late Permian

A therocephalian with a short skull.

Ruthenosaurus[161]

Gen. et sp.

Valid

Upper Sakmarian - Lower Lopingian

Grès Rouge Group

A large caseid. The type species is Ruthenosaurus russellorum

Syops[157]

Gen. et comb.

Valid

  • Kammerer
  • Angielczyk
  • Fröbisch

Late Permian

A dicynodont. A new genus for "Dicynodon" vanhoepeni.

Tiarajudens[162]

Gen. et sp.

Valid

Capitanian

Rio do Rasto Formation

A basal, saber-toothed herbivorous anomodont. The type species is Tiarajudens eccentricus.

Newly named non-eutherian mammals

Name Novelty Status Authors Age Unit Location Notes Images

Cronopio[163]

Gen. et sp.

Valid

  • Rougier
  • Apesteguía
  • Gaetano

Cenomanian

Candeleros Formation

A "saber-toothed" dryolestoid.

Crusafontia amoae[164]

Species

Valid

Hauterivian to Barremian

A dryolestidan mammal.

Iberica[165]

Gen. et sp.

Valid

  • Badiola
  • Canudo
  • Cuenca-Bescós

Hauterivian/Barremian

Galve

A eobaatarid or a possible plagiaulacid multituberculate.

Invictokoala[166]

Gen. et sp.

Valid

  • Price
  • Hocknull

Middle Pleistocene

A Koala.

Liaoconodon[167]

Gen. et sp.

Valid

  • Meng
  • Wang
  • Li

Aptian

Jiufotang Formation

An "eutriconodont".

Naraboryctes[168]

Gen. et sp.

Valid

Early Miocene

A marsupial mole.

Oklatheridium minax[169]

Species

Valid

  • Davis
  • Cifelli

Aptian-Albian

Antlers Formation

A deltatheroidan mammal.

Sineleutherus issedonicus[170]

Species

Valid

  • Averianov
  • Lopatin
  • Krasnolutskii

Bathonian

Itat Formation

A haramiyidan allotherian.

Newly named eutherians

Name Novelty Status Authors Age Unit Location Notes Images

Adelpharctos ginsburgi[171]

Sp. nov

Valid

  • de Bonis

Late Oligocene

A hemicyonine bear, a species of Adelpharctos.

Aegyptocetus[172]

Gen. et sp.

Valid

  • Bianucci
  • Gingerich

Middle Eocene

Gebel Hof Formation

A protocetid cetacean.

Agriarctos beatrix[173]

Sp. nov

Valid

  • Abella, Montoya & Morales

Middle Miocene

Calatayud-Daroca Basin

An ailuropodine bear, a species of Agriarctos.

Antarctodon[174]

Gen. et sp.

Valid

Late Ypresian

La Meseta Formation

An astrapotherian.

Asilifelis[175]

Gen. et sp. nov

Valid

  • Werdelin

Lower Miocene

Hiwegi Formation

A felid. The type species is Asilifelis coteae.

Betonnia[176]

Gen. et sp.

Valid

  • Williamson
  • Weil
  • Standhardt

Middle/Late Puercan

Nacimiento Formation

A cimolestid cimolesta.

Balochititanops[177]

Gen. et sp.

Valid

  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid.

Caviodon cuyano[178]

Species

Valid

Late Miocene or Pliocene.

Aisol Formation

A cardiomyine rodent, a species of Caviodon.

Chacomylus[179]

Gen. et sp.

Valid

  • Williamson
  • Weil

Paleocene

Nacimiento Formation

A hyopsodontid condylarth.

Chacopterygus[176]

Gen. et sp.

Valid

  • Williamson
  • Weil
  • Standhardt

Middle Puercan

Nacimiento Formation

A cimolestid cimolesta.

Coelodonta thibetana[180]

Species

Valid

Middle Pliocene

Woolly rhinoceros.

Comahuetherium[181]

Gen. et sp.

Valid

  • Kramarz
  • Bond

Colhuehuapian

Cerro Bandera Formation

An astrapotheriid.

Cynthiacetus peruvianus[182]

Species

Valid

  • Martínez-Cáceres
  • Muizon

Late Eocene-Early Oligocene

Otuma Formation

A basilosaurid cetacean.

Delotrochanter[183]

Gen. et 3 sp. nov.

Valid

  • Hunt

Arikareean

A temnocyonine. The type species is Delotrochanter oryktes; genus also contains D. petersoni and D. major

Diplacodon gigan[184]

Species

Valid

  • Mihlbachler

Middle Eocene

Wiggins Formation

A brontotheriid.

Eotitanops pakistanensis[177]

Species

Valid

  • Missiaen
  • Gunnell
  • Gingerich

Ypresian

Ghazij Formation

A brontotheriid.

Griphotherion[185]

Gen. et sp.

Valid

  • López
  • Powell

Eocene

A rodent-like notoungulate.

Hiskatherium[186]

Gen. et sp.

Valid

  • Pujos
  • Iuliis
  • Quispe

Santacrucian

A small ground sloth.

Hystrix paukensis[187]

Species

Valid

  • Yuichiro Nishioka
  • Zin-Maung-Maung-Thein
  • et al.[CAL 65]
late Miocene/early Pliocene new species in the genus Hystrix (Old World porcupine).

Juramaia[188]

Gen. et sp.

Valid

Callovian to Bathonian

Tiaojishan Formation

A basal eutherian. The type species is Juramaia sinensis.

Karydomys debruijni[189]

Species

Valid

  • Maridet et al.

Early Miocene

A cricetid, a species of Karydomys.

Khoratpithecus ayeyarwadyensis[190]

Species

Valid

Late Miocene

Irrawaddy Formation

A hominoid.

Kuntinaru[191]

Gen. et sp.

Valid

Deseadan

A dasypodid armadillo.

Lycophocyon[192]

Gen. et sp.

Valid

  • Tomiya

?Uintan-Duchesnean

Santiago Formation

A caniformian carnivoran.

Mammacyon ferocior[183]

Sp. nov.

Valid

  • Hunt

Mid- or early late Arikareean

Arikaree Group

A temnocyonine, a species of Mammacyon

Martes ginsburgi[193]

Sp. nov

Valid

  • Montoya, Morales & Abella

Upper Miocene

A marten.

Megacricetodon beijiangensis[189]

Species

Valid

  • Maridet et al.

Early Miocene

A cricetid, a species of Megacricetodon.

Mescalerolemur[194]

Gen. et sp.

Valid

  • Kirk
  • Williams

Uintan

Devil’s Graveyard Formation

A adapiform strepsirrhini.

Mesolambdolophus[195]

Gen. et sp.

Valid

  • Holbrook
  • Lapergola

Bridgerian North American Stage

Bridger Formation

A small odd-toed ungulate of uncertain phylogenetic position, possibly related to Tapiromorpha. The type species is Mesolambdolophus setoni.

Mionothropus[196]

Gen. et sp.

Valid

  • De Iuliis
  • Gaudin
  • Vicars

Huayquerian

A megatheriid sloth.

Mondegodon[197]

Gen. et sp.

Valid

  • Tabuce
  • Clavel
  • Antunes

Earliest Eocene

A mesonychian.

Nuralagus[198]

Gen. et sp.

Valid

  • Quintana
  • Köhler
  • Moyà-Solà

Early Pliocene

A leporid rabbit.

Ocucajea[199]

Gen. et sp.

Valid

Bartonian

Paracas Formation

A basilosaurid cetacean.

Oldrichpedetes[200]

Gen. et comb. et 3 sp. nov

Valid

  • Pickford & Mein

Early Miocene to early Pliocene

A pedetid rodent, a new genus for "Megapedetes" pickfordi (Mein & Senut, 2003). Genus also contains 3 new species: Oldrichpedetes fejfari, O. brigitteae and O. praecursor.

Phoxomylus[201]

Gen. et sp.

Valid

  • Fox

Early Tiffanian

Paskapoo Formation

A palaechthonid plesiadapiform.

Protoryx tuvaensis[202]

Species

Valid

  • Dmitrieva
  • Serdyuk

Late Miocene

A bovid.

Protungulatum coombsi[203]

Species

Valid

Maastrichtian

Hell Creek Formation

A species of Protungulatum.

Rudiocyon[183]

Gen. et sp. nov.

Valid

  • Hunt

Arikareean

John Day Formation

A temnocyonine. The type species is Rudiocyon amplidens

Rusingapedetes[200]

Gen. et sp. nov

Valid

  • Pickford & Mein

Early Miocene

A pedetid rodent. The type species is Rusingapedetes tsujikawai.

Rytiodus heali[204]

Species

Valid

  • Domning
  • Sorbi

Miocene

A dugongid sirenian, a species of Rytiodus.

Supayacetus[199]

Gen. et sp.

Valid

Bartonian

Paracas Formation

A basilosaurid cetacean.

Temnocyon fingeruti[183]

Sp. nov.

Valid

  • Hunt

Latest Oligocene, late mid-Arikareean

John Day Formation

A temnocyonine, a species of Temnocyon

Temnocyon macrogenys[183]

Sp. nov.

Valid

  • Hunt

Latest Arikareean

Anderson Ranch Formation

A temnocyonine, a species of Temnocyon

Temnocyon subferox[183]

Sp. nov.

Valid

  • Hunt

Early Arikareean

John Day Formation

A temnocyonine, a species of Temnocyon

Turiasorex[205]

Gen. et sp.

Valid

  • van Dam
  • van den Hoek Ostende
  • Reumer

Middle - Late Miocene

A short-snouted shrew.

Umayodus[206]

Gen. et sp.

Valid

  • Gelfo
  • Sigé

Late Paleocene/Earliest Eocene

Muñami Formation

A didolodontid "condylarth".

Plants

Angiosperms

Name Novelty Status Authors Age Unit Location Notes Images

Palmoxylon lametaei[207]

Species

Valid

  • Dutta
  • Ambwani
  • Estrada-Ruiz

Maastrichtian

Lameta Formation

Leefructus[208]

Gen. et sp.

Valid

Early Cretaceous

Yixian Formation

A basal eudicot

Gymnosperms

Name Novelty Status Authors Age Unit Location Notes Images

Qataniaria[209]

Gen. et sp.

Valid

  • Krassilov
  • Schrank

Albian

Hatira Formation

Pteridophytes

Name Novelty Status Authors Age Unit Location Notes Images

Equisetum thermale[210]

sp. nov

Valid

Channing, Zamuner, Edwards, & Guido

Callovian - Tithonian

La Matilde Formation

 Argentina

oldest crown group Equisetum species

Footnotes

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.

  1. ^ Liu, Steiner, Dunlop, Keupp, Shu, Ou, Han, Zang, Zang.
  2. ^ Gallego, Rébori, Zavattieri, Sinitshenkova, Lara and Martins-Neto.
  3. ^ Gai, Donoghue, Zhu, Janvier, Stampanoni.
  4. ^ Underwood, Goswami Prasad, Verma and Flynn
  5. ^ Fischer, Voigt, Schneider, Buchwitz and Voigt
  6. ^ Pradel, Tafforeau, Maisey, Janvier.
  7. ^ Downs, Daeschler, Jenkins and Shubin.
  8. ^ Wu, Sun, Xu, Hao, Jiang, Sun.
  9. ^ Henrici, Martens, Berman, Sumida.
  10. ^ Berman, Henrici, Martens, Sumida, Anderson
  11. ^ a b Reisz, Liu, Li, Müller.
  12. ^ Gaffney, Meylan, Wood, Simons and Campos
  13. ^ Slater, Reolid, Wood, Schouten and Benton
  14. ^ Knauss, Joyce, Lyson and Pearson
  15. ^ de la Fuente, Umazano, Sterli and Carballido
  16. ^ Danilov, Zvonok, Syromyatnikova and Udovichenko
  17. ^ Fischer, Masure, Arkhangelsky, Godefroit.
  18. ^ Liu, Rieppel, Jiang, Aitchison, Motani, Zhang, Zhou, Sun.
  19. ^ a b Benson, Ketchum, Noè, Gómez-Pérez.
  20. ^ Müller, Hipsley, Head, Kardjilov, Hilger, Wuttke, Reisz.
  21. ^ a b Carvalho, Teixeira, Ferraz, Ribeiro, Martinelli, Neto, Sertich, Cunha, Cunha, Ferraz.
  22. ^ Kellner, Campos, Riff, de Andrade.
  23. ^ a b de Andrade, Edmonds, Benton, Schouten.
  24. ^ Buscalioni, Piras, Vullo, Signore, Barbera
  25. ^ Lauprasert, Laojumpon, Saenphala, Cuny, Thirakhupt, Suteethorn
  26. ^ Hu, Li, Hou, Xu.
  27. ^ Bocheński, Tomek, Bujoczek, Wertz.
  28. ^ Mourer-Chauviré, Tabuce, Mahboubi, Adaciy, Bensalah.
  29. ^ Kurochkin, Zelenkov, Averianov, Leshchinskiy.
  30. ^ Ji, Atterholt, O'Connor, Lamanna, Harrs, Li, You, Dodson.
  31. ^ Gates, Horner, Hanna, Nelson.
  32. ^ Nesbitt, Clarke, Turner, Norell.
  33. ^ Mateus, Jacobs, Schulp, Polcyn, Tavares, Neto, Morais, Antunes.
  34. ^ Kellner, Rubilar-Rogers, Vargas, Suárez
  35. ^ Sues, Nesbitt, Berman, Henrici
  36. ^ Fernández-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola, Salgado.
  37. ^ Martinez, Sereno, Alcober, Colombi, Renne, Montañez, Currie.
  38. ^ Makovicky, Kilbourne, Sadleir, Norell.
  39. ^ a b Novas, Ezcurra, Chatterjee, Kutty.
  40. ^ Huh, Lee, Kim, Lim, Godefroit.
  41. ^ Apaldetti, Martinez, Alcober, Pol.
  42. ^ Xu, Sullivan, Pittman, Choiniere, Hone, Upchurch, Tan, Xiao, Tan, Han.
  43. ^ Xu, Tan, Sullivan, Han, Xiao.
  44. ^ Jiang, Li, Peng, Ye
  45. ^ Kellner, Azvedo, Machado, Carvalho, Henriques.
  46. ^ Schultz, Bittencourt, Soares, Fortier, Silva, Langer.
  47. ^ Filippi, Canudo, Salgado, Garrido, Garcia, Cerda, Otero.
  48. ^ Xu, Kobayashi, Lü, Lee, Liu, Tanaka, Zhang, Jia, Zhang.
  49. ^ Farke, Ryan, Barrett, Tanke, Braman, Loewen, Graham.
  50. ^ Zanno, Varricchio, O'Connor, Titus, Knell.
  51. ^ Zaher, Pol, Carvalho, Nascimento, Riccomini, Larson, Juarez-Valieri, Pires-Domingues, da Silva Jr., Campos.
  52. ^ Carr, Williamson, Britt, Stadtman.
  53. ^ Xu, You, Du, Han.
  54. ^ Hone, Wang, Sullivan, Zhao, Chen, Li, Ji, Ji, Xu.
  55. ^ Lü, Xu, Chang, Zhang.
  56. ^ De Oliveira, Schultz, Soares, Rodrigues.
  57. ^ Fröbisch, Schoch, Müller, Schindler, Schweiss.
  58. ^ a b Reisz, Maddin, Fröbisch, Falconnet.
  59. ^ Cisneros, Abdala, Rubidge, Dentzien-Dias, Bueno.
  60. ^ Cuenca-Bescós, Badiola, Canudo, Gasca and Moreno-Azanza.
  61. ^ Archer, Beck, Gott, Hand, Godthelp, Black.
  62. ^ Bond, Kramarz, MacPhee, Reguero.
  63. ^ Vucetich, Deschamps, Morgan and Forasiepi.
  64. ^ Deng, Wang, Fortelius, Li, Wang, Tseng, Takeuchi, Saylor, Säilä, Xie
  65. ^ Naoko Egi, Takehisa Tsubamoto, Takeshi Nishimura, Tsuyoshi Ito, Thaung-Htike and Masanaru Takai.
  66. ^ Luo, Yuan, Meng, Ji.
  67. ^ Jaeger, Soe, Chavasseau, Coster, Emonet, Guy, Lebrum, Maung, Shwe, Tun, Rugbumrung, Bocherens, Benammi, Chaivanich, Tafforeau, Chaimanee.
  68. ^ Billet, Hautier, de Muizon, Valentin.
  69. ^ a b Uhen, Pyenson, Devries, Urbina, Renne.
  70. ^ Archibald, Zhang, Harper and Cifelli
  71. ^ Sun, Dilcher, Wang, Chen.

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