Angular angel shark

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Angular angel shark
Conservation status
Endangered (IUCN 3.1)
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Chondrichthyes
Subclass:	Elasmobranchii
Order:	Squatiniformes
Family:	Squatinidae
Genus:	Squatina
Species:	S. guggenheim
Binomial name
*Squatina guggenheim* Marini, 1936

Range of Angular angelshark (in blue)

Sharks portal

The angular angel shark, Squatina guggenheim, is an angel shark of the family Squatinidae found off southern Brazil and Argentina at depths of between 4 and 265 m. Its length is up to 1.3 m.

Reproduction is ovoviviparous, with 6 to 8 pups in a litter.

It was formerly named Squatina occulta.

References

Froese, Rainer, and Daniel Pauly, eds. (2006). "Squatina guggenheim" in FishBase. July 2006 version.
Compagno, Dando, & Fowler, Sharks of the World, Princeton University Press, New Jersey 2005 ISBN 0-691-12072-2

Extant shark species

Kingdom Animalia · Phylum Chordata · Subphylum Vertebrata · Class Chondrichthyes · Subclass Elasmobranchii

Order Hexanchiformes

Chlamydoselachidae

Chlamydoselachus

Frilled shark (C. anguineus) · Southern African frilled shark (C. africana)

Hexanchidae
(Cow sharks)

Heptranchias	Sharpnose sevengill shark (H. perlo)

Hexanchus	Bluntnose sixgill shark (H. griseus) · Bigeyed sixgill shark (H. nakamurai)

Notorynchus	Broadnose sevengill shark (N. cepedianus)

Order Squaliformes

Centrophoridae
(Gulper sharks)

Centrophorus	Needle dogfish (C. acus) · Dwarf gulper shark (C. atromarginatus) · Gulper shark (C. granulosus) · Dumb gulper shark (C. harrissoni) · Blackfin gulper shark (C. isodon) · Lowfin gulper shark (C. lusitanicus) · Smallfin gulper shark (C. moluccensis) · Taiwan gulper shark (C. niaukang) · Leafscale gulper shark (C. squamosus) · Mosaic gulper shark (C. tessellatus) · Little gulper shark (C. uyato)

Deania	Birdbeak dogfish (D. calcea) · Rough longnose dogfish (D. hystricosa) · Arrowhead dogfish (D. profundorum) · Longsnout dogfish (D. quadrispinosum)

Dalatiidae

Euprotomicroides	Taillight shark (E. zantedeschia)

Heteroscymnoides	Longnose pygmy shark (H. marleyi)

Mollisquama	Pocket shark (M. parini)

Dalatias	Kitefin shark (D. licha)

Isistius	Cookiecutter shark (I. brasiliensis) · South China cookiecutter shark (I. labialis) · Largetooth cookiecutter shark (I. plutodus)

Euprotomicrus	Pygmy shark (E. bispinatus)

Squaliolus	Smalleye pygmy shark (S. aliae) · Spined pygmy shark (S. laticaudus)

Echinorhinidae

Echinorhinus

Bramble shark (E. brucus) · Prickly shark (E. cookei)

Etmopteridae

Aculeola	Hooktooth dogfish (A. nigra)

Centroscyllium	Highfin dogfish (C. excelsum) · Black dogfish (C. fabricii) · Granular dogfish (C. granulatum) · Bareskin dogfish (C. kamoharai) · Combtooth dogfish (C. nigrum) · Ornate dogfish (C. ornatum) · Whitefin dogfish (C. ritteri)

Etmopterus (Lantern sharks)	New Zealand lanternshark (E. baxteri) · Blurred lanternshark (E. bigelowi) · Shorttail lanternshark (E. brachyurus) · Lined lanternshark (E. bullisi) · E. burgessi · Cylindrical lanternshark (E. carteri) · Tailspot lanternshark (E. caudistigmus) · Combtooth lanternshark (E. decacuspidatus) · Pink lanternshark (E. dianthus) · E. dislineatus · Blackmouth lanternshark (E. evansi) · Pygmy lanternshark (E. fusus) · Broadbanded lanternshark (E. gracilispinis) · Southern lanternshark (E. granulosus) · Caribbean lanternshark (E. hillianus) · Smalleye lantern shark (E. litvinovi) · Blackbelly lanternshark (E. lucifer) · Slendertail lanternshark (E. molleri) · Dwarf lanternshark (E. perryi) · African lanternshark (E. polli) · Great lanternshark (E. princeps) · False lanternshark (E. pseudosqualiolus) · Smooth lanternshark (E. pusillus) · Dense-scale lantern shark (E. pycnolepis) · West Indian lanternshark (E. robinsi) · Fringefin lanternshark (E. schultzi) · Thorny lanternshark (E. sentosus) · Velvet belly lantern shark (E. spinax) · Splendid lanternshark (E. splendidus) · Tasmanian lanternshark (E. tasmaniensis) · Brown lanternshark (E. unicolor) · Hawaiian lanternshark (E. villosus) · Green lanternshark (E. virens)

Miroscyllium	Rasptooth dogfish (M. sheikoi)

Trigonognathus	Viper dogfish (T. kabeyai)

Oxynotidae
(Rough sharks)

Oxynotus

Prickly dogfish (O. bruniensis) · Caribbean roughshark (O. caribbaeus) · Angular roughshark (O. centrina) · Japanese roughshark (O. japonicus) · Sailfin roughshark (O. paradoxus)

Somniosidae
(Sleeper sharks)

Squalidae
(Dogfish sharks)

Cirrhigaleus	Roughskin spurdog (C. asper) · Mandarin dogfish (C. barbifer)

Squalus (Spurdogs)	Spiny dogfish (S. acanthias) · Eastern highfin spurdog (S. albifrons) · S. acutirostris · Western highfin spurdog (S. altipinnis) · Longnose spurdog (S. blainville) · Fatspine spurdog (S. crassispinus) · Cuban dogfish (S. cubensis) · Eastern longnose spurdog (S. grahami) · Japanese spurdog (S. japonicus) · Shortnose spurdog (S. megalops) · Blacktailed spurdog (S. melanurus) · Shortspine spurdog (S. mitsukurii) · Bartail spurdog (S. notocaudatus) · Western longnose spurdog (S. nasutus) · Cyrano spurdog (S. rancureli) · Squalus suckleyi

Order Pristiophoriformes (Sawsharks)

Pristiophoridae

Pliotrema	Sixgill sawshark (P. warreni)

Pristiophorus	Longnose sawshark (P. cirratus) · Tropical sawshark (P. delicatus) · Japanese sawshark (P. japonicus) · Shortnose sawshark (P. nudipinnis) · Bahamas sawshark (P. schroederi) · Eastern Australian sawshark (Pristiophorus peroniensis) · Philippine sawshark (Pristiophorus sp. C) · Dwarf sawshark (Pristiophorus sp. D)

Order Squatiniformes (Angel sharks)

Squatinidae

Squatina

Sawback angelshark (S. aculeata) · African angelshark (S. africana) · Eastern Australian angelshark (Squatina albipunctata) · Argentine angelshark (S. argentina) · Chilean angelshark (S. armata) · Australian angelshark (S. australis) · Pacific angelshark (S. californica) · Sand devil (S. dumeril) · Taiwan angelshark (S. formosa) · Angular angel shark (S. guggenheim) · S. heteroptera · Japanese angelshark (S. japonica) · Indonesian angelshark (S. legnota) · Cortez angelshark (S. mexicana) · Clouded angelshark (S. nebulosa) · Smoothback angelshark (S. oculata) · S. punctata · Western Australian angelshark (Squatina pseudocellata) · Angelshark (S. squatina) · Ornate angelshark (S. tergocellata) · Ocellated angelshark (S. tergocellatoides)

Order Heterodontiformes (Bullhead sharks)

Heterodontidae

Heterodontus

Horn shark (H. francisci) · Crested bullhead shark (H. galeatus) · Japanese bullhead shark (H. japonicus) · Mexican hornshark (H. mexicanus) · Oman bullhead shark (H. omanensis) · Port Jackson shark (H. portusjacksoni) · Galapagos bullhead shark (H. quoyi) · Whitespotted bullhead shark (H. ramalheira) · Zebra bullhead shark (H. zebra)

Order Orectolobiformes (Carpet sharks)

Brachaeluridae

Brachaelurus	Blind shark (B. waddi)

Heteroscyllium	Bluegrey carpetshark (H. colcloughi)

Ginglymostomatidae
(Nurse sharks)

Ginglymostoma	Nurse shark (G. cirratum)

Nebrius	Tawny nurse shark (N. ferrugineus)

Pseudoginglymostoma	Short-tail nurse shark (P. brevicaudatum)

Hemiscylliidae
(Bamboo sharks)

Chiloscyllium	Arabian carpetshark (C. arabicum) · Burmese bamboo shark (C. burmensis) · Bluespotted bamboo shark (C. caerulopunctatum) · Grey bamboo shark (C. griseum) · Hasselt's bamboo shark (C. hasseltii) · Slender bamboo shark (C. indicum) · Whitespotted bamboo shark (C. plagiosum) · Brownbanded bamboo shark (C. punctatum)

Hemiscyllium	Indonesian speckled carpetshark (H. freycineti) · H. galei · Papuan epaulette shark (H. hallstromi) · H. henryi · Epaulette shark (H. ocellatum) · Hooded carpetshark (H. strahani) · Speckled carpetshark (H. trispeculare)

Orectolobidae
(Wobbegongs)

Eucrossorhinus	Tasselled wobbegong (E. dasypogon)

Orectolobus	Floral banded wobbegong (O. floridus) · Banded wobbegong (O. halei) · Western wobbegong (O. hutchinsi) · Japanese wobbegong (O. japonicus) · Spotted wobbegong (O. maculatus) · Ornate wobbegong (O. ornatus) · Dwarf spotted wobbegong (O. parvimaculatus) · Network wobbegong (O. reticulatus) · Northern wobbegong (O. wardi)

Sutorectus	Cobbler wobbegong (S. tentaculatus)

Parascylliidae
(Collared carpet sharks)

Cirrhoscyllium	Barbelthroat carpetshark (C. expolitum) · Taiwan saddled carpetshark (C. formosanum) · Saddle carpetshark (C. japonicum)

Parascyllium	Collared carpetshark (P. collare) · Rusty carpetshark (P. ferrugineum) · Ginger carpetshark (P. sparsimaculatum) · Necklace carpetshark (P. variolatum)

Rhincodontidae

Rhincodon

Whale shark (R. typus)

Stegostomatidae

Stegostoma

Zebra shark (S. fasciatum)

Order Carcharhiniformes (Ground sharks)

Hemigaleidae
(Weasel sharks)

Hemipristis	Snaggletooth shark (H. elongata)

Chaenogaleus	Hooktooth shark (C. macrostoma)

Hemigaleus	Australian weasel shark (H. australiensis) · Sicklefin weasel shark (H. microstoma)

Paragaleus	Whitetip weasel shark (P. leucolomatus) · Atlantic weasel shark (P. pectoralis) · Slender weasel shark (P. randalli) · Straight-tooth weasel shark (P. tengi)

Leptochariidae

Leptocharias

Barbeled houndshark (L. smithii)

Proscylliidae
(Finback sharks)

Ctenacis	Harlequin catshark (C. fehlmanni)

Eridacnis	Cuban ribbontail catshark (E. barbouri) · Pygmy ribbontail catshark (E. radcliffei) · African ribbontail catshark (E. sinuans)

Proscyllium	Graceful catshark (P. habereri) · P. venustum

Pseudotriakidae

Gollum	Slender smooth-hound (G. attenuatus)

Pseudotriakis	False catshark (P. microdon)

Sphyrnidae
(Hammerhead sharks)

Eusphyra	Winghead shark (E. blochii)

Sphyrna	Scalloped bonnethead (S. corona) · Whitefin hammerhead (S. couardi) · Scalloped hammerhead (S. lewini) · Scoophead (S. media) · Great hammerhead (S. mokarran) · Bonnethead (S. tiburo) · Smalleye hammerhead (S. tudes) · Smooth hammerhead (S. zygaena)

Triakidae
(Houndsharks)

Furgaleus	Whiskery shark (F. macki)

Galeorhinus	School shark (G. galeus)

Gogolia	Sailback houndshark (G. filewoodi)

Hemitriakis	Deepwater sicklefin hound shark (H. abdita) · Sicklefin hound shark (H. falcata) · Japanese topeshark (H. japanica) · Whitefin topeshark (H. leucoperiptera) · Ocelate topeshark (Hemitriakis Sp.A)

Hypogaleus	Blacktip tope (H. hyugaensis)

Iago	Longnose houndshark (I. garricki) · Bigeye houndshark (I. omanensis) · Lowfin houndshark (Iago Sp.A)

Mustelus (Smooth-hounds)	M. albipinnis · Gummy shark (M. antarcticus) · Starry smooth-hound (M. asterias) · Grey smooth-hound (M. californicus) · Dusky smooth-hound (M. canis) · Sharptooth smooth-hound (M. dorsalis) · Striped smooth-hound (M. fasciatus) · Spotless smooth-hound (M. griseus) · Brown smooth-hound (M. henlei) · Smalleye smooth-hound (M. higmani) · Spotted estuary smooth-hound (M. lenticulatus) · Sicklefin smooth-hound (M. lunulatus) · Starspotted smooth-hound (M. manazo) · Speckled smooth-hound (M. mento) · M. minicanis · Arabian smooth-hound (M. mosis) · Common smooth-hound (M. mustelus) · Narrowfin smooth-hound (M. norrisi) · Whitespotted smooth-hound (M. palumbes) · Blackspotted smooth-hound (M. punctulatus) · M. ravidus · Narrownose smooth-hound (M. schmitti) · Gulf smoothhound (M. sinusmexicanus) · Humpback smooth-hound (M. whitneyi) · M. widodoi

Scylliogaleus	Flapnose houndshark (S. quecketti)

Triakis	Sharpfin houndshark (T. acutipinna) · Spotted houndshark (T. maculata) · Sharptooth houndshark (T. megalopterus) · Banded houndshark (T. scyllium) · Leopard shark (T. semifasciata)

Carcharhinidae

Large family listed below

Scyliorhinidae

Large family listed below

Family Carcharhinidae (Requiem sharks)

Carcharhinus	Blacknose shark (C. acronotus) · Silvertip shark (C. albimarginatus) · Bignose shark (C. altimus) · Graceful shark (C. amblyrhynchoides) · Grey reef shark (C. amblyrhynchos) · Pigeye shark (C. amboinensis) · Borneo shark (C. borneensis) · Copper shark (C. brachyurus) · Spinner shark (C. brevipinna) · Nervous shark (C. cautus) · Whitecheek shark (C. dussumieri) · Silky shark (C. falciformis) · Creek whaler (C. fitzroyensis) · Galapagos shark (C. galapagenisis) · Pondicherry shark (C. hemiodon) · Finetooth shark (C. isodon) · Smoothtooth blacktip shark (C. leiodon) · Bull shark (C. leucas) · Blacktip shark (C. limbatus) · Oceanic whitetip shark (C. longimanus) · Hardnose shark (C. macloti) · Blacktip reef shark (C. melanopterus) · Dusky shark (C. obscurus) · Caribbean reef shark (C. perezii) · Sandbar shark (C. plumbeus) · Smalltail shark (C. porosus) · Blackspot shark (C. sealei) · Night shark (C. signatus) · Spottail shark (C. sorrah) · Australian blacktip shark (C. tilstoni)

Galeocerdo	Tiger shark (G. cuvier)

Glyphis (River sharks)	Ganges shark (G. gangeticus) · Northern river shark (G. garricki) · Speartooth shark (G. glyphis) · Irrawaddy river shark (G. siamensis) · Borneo river shark (Glyphis sp. B)

Isogomphodon	Daggernose shark (I. oxyrhynchus)

Lamiopsis	Broadfin shark (L. temminckii)

Loxodon	Sliteye shark (L. macrorhinus)

Nasolamia	Whitenose shark (N. velox)

Negaprion	Sicklefin lemon shark (N. acutidens) · Lemon shark (N. brevirostris)

Prionace	Blue shark (P. glauca)

Rhizoprionodon	Milk shark (R. acutus) · Brazilian sharpnose shark (R. lalandii) · Pacific sharpnose shark (R. longurio) · Grey sharpnose shark (R. oligolinx) · Caribbean sharpnose shark (R. porosus) · Australian sharpnose shark (R. taylori) · Atlantic sharpnose shark (R. terraenovae)

Scoliodon	Spadenose shark (S. laticaudus)

Triaenodon	Whitetip reef shark (T. obesus)

Family Scyliorhinidae (Catsharks)

Apristurus	Flatnose cat shark (A. acanutus) · A. albisoma · A. aphyodes · Atlantic ghost catshark (A. atlanticus) · Brown catshark (A. brunneus) · Hoary catshark (A. canutus) · Flaccid catshark (A. exsanguis) · A. fedorovi · Humpback cat shark (A. gibbosus) · Longfin catshark (A. herklotsi) · Smallbelly catshark (A. indicus) · A. internatus · Broadnose catshark (A. investigatoris) · Japanese catshark (A. japonicus) · Longnose catshark (A. kampae) · Iceland catshark (A. laurussonii) · Longhead catshark (A. longicephalus) · Flathead catshark (A. macrorhynchus) · Broadmouth cat shark (A. macrostomus) · Ghost catshark (A. manis) · Black roughscale catshark (A. melanoasper) · Smalleye catshark (A. microps) · Smalldorsal cat shark (A. micropterygeus) · Largenose catshark (A. nasutus) · Smallfin catshark (A. parvipinnis) · A. pinguis · Spatulasnout catshark (A. platyrhynchus) · Deepwater catshark (A. profundorum) · Broadgill catshark (A. riveri) · Saldanha catshark (A. saldanha) · Pale catshark (A. sibogae) · South China catshark (A. sinensis) · Spongehead catshark (A. spongiceps) · Panama ghost catshark (A. stenseni)

Asymbolus	Australian spotted catshark (A. analis) · A. funebris · Western spotted catshark (A. occiduus) · Pale spotted catshark (A. pallidus) · A. parvus · A. rubiginosus · Variegated catshark (A. submaculatus) · Gulf catshark (A. vincenti)

Atelomycterus	A. baliensis · Banded sand catshark (A. fasciatus) · Australian marbled catshark (A. macleayi) · Coral catshark (A. marmoratus)

Aulohalaelurus	New Caledonia catshark (A. kanakorum) · Australian blackspotted catshark (A. labiosus)

Cephaloscyllium	Whitefin swellshark (C. albipinnum) · Circle-blotch pygmy swellshark (C. circulopullum) · Cook's swellshark (C. cooki) · Reticulated swellshark (C. fasciatum) · Australian reticulate swellshark (C. hicosellum) · Draughtsboard shark (C. isabellum) · Australian swellshark (C. laticeps) · Spotted swellshark (C. maculatum) · Leopard-spotted swellshark (C. pardelotum) · Painted swellshark (C. pictum) · Sarawak pygmy swellshark (C. sarawakensis) · Flagtail swellshark (C. signourum) · Indian swellshark (C. silasi) · Speckled swellshark (C. speccum) · Balloon shark (C. sufflans) · Blotchy swellshark (C. umbratile) · Saddled swellshark (C. variegatum) · Swellshark (C. ventriosum) · Narrowbar swellshark (C. zebrum)

Cephalurus	Lollipop catshark (C. cephalus)

Figaro	Australian sawtail catshark (F. boardmani) · Northern sawtail catshark (F. striatus)

Galeus	Antilles catshark (G. antillensis) · Roughtail catshark (G. arae) · Atlantic sawtail cat shark (G. atlanticus) · Longfin sawtail cat shark (G. cadenati) · Gecko catshark (G. eastmani) · Slender sawtail catshark (G. gracilis) · Longnose sawtail cat shark (G. longirostris) · Blackmouth catshark (G. melastomus) · Southern sawtail catshark (G. mincaronei) · Mouse catshark (G. murinus) · Broadfin sawtail catshark (G. nipponensis) · Peppered catshark (G. piperatus) · African sawtail catshark (G. polli) · G. priapus · Blacktip sawtail catshark (G. sauteri) · Dwarf sawtail catshark (G. schultzi) · Springer's sawtail cat shark (G. springeri)

Halaelurus	Arabian catshark (H. alcockii) · Speckled catshark (H. boesemani) · Blackspotted catshark (H. buergeri) · Dusky catshark (H. canescens) · Broadhead cat shark (H. clevai) · New Zealand catshark (H. dawsoni) · Bristly catshark (H. hispidus) · Spotless catshark (H. immaculatus) · Lined catshark (H. lineatus) · Mud catshark (H. lutarius) · Tiger catshark (H. natalensis) · Quagga catshark (H. quagga)

Haploblepharus	Puffadder shyshark (H. edwardsii) · Brown shyshark (H. fuscus) · Natal shyshark (H. kistnasamyi) · Dark shyshark (H. pictus)

Holohalaelurus	H. favus · H. grennian · Crying izak (H. melanostigma) · African spotted catshark (H. punctatus) · Izak catshark (H. regani)

Parmaturus	White-tip catshark (P. albimarginatus) · White-clasper catshark (P. albipenis) · Beige catshark (P. bigus) · Campeche catshark (P. campechiensis) · Velvet catshark (P. lanatus) · McMillan's catshark (P. macmillani) · Blackgill catshark (P. melanobranchus) · Salamander shark (P. pilosus) · Filetail catshark (P. xaniurus) · Shorttail catshark (Parmaturus sp. A)

Pentanchus	Onefin catshark (P. profundicolus)

Poroderma	Pyjama catshark (P. africanum) · Leopard catshark (P. pantherinum)

Schroederichthys	Narrowmouthed catshark (S. bivius) · Redspotted catshark (S. chilensis) · Narrowtail catshark (S. maculatus) · Lizard catshark (S. saurisqualus) · Slender catshark (S. tenuis)

Scyliorhinus	Polkadot catshark (S. besnardi) · Boa catshark (S. boa) · Small-spotted catshark (S. canicula) · Yellowspotted catshark (S. capensis) · West African catshark (S. cervigoni) · Comoro cat shark (S. comoroensis) · Brownspotted catshark (S. garmani) · Freckled catshark (S. haeckelii) · Whitesaddled catshark (S. hesperius) · Blotched catshark (S. meadi) · Chain catshark (S. retifer) · Nursehound (S. stellaris) · Izu cat shark (S. tokubee) · Cloudy catshark (S. torazame) · Dwarf catshark (S. torrei)

Order Lamniformes (Mackerel sharks)

Alopiidae

Alopias
(Thresher sharks)

Pelagic thresher (A. pelagicus) · Bigeye thresher (A. superciliosus) · Common thresher (A. vulpinus)

Cetorhinidae

Cetorhinus

Basking shark (C. maximus)

Lamnidae

Carcharodon	Great white shark (C. carcharias)

Isurus	Shortfin mako shark (I. oxyrinchus) · Longfin mako shark (I. paucus)

Lamna	Salmon shark (L. ditropis) · Porbeagle (L. nasus)

Megachasmidae

Megachasma

Megamouth shark (M. pelagios)

Mitsukurinidae

Mitsukurina

Goblin shark (M. owstoni)

Odontaspididae

Carcharias	Grey nurse shark (C. taurus) · Indian sand tiger (C. tricuspidatus)

Odontaspis	Smalltooth sand tiger (O. ferox) · Bigeye sand tiger (O. noronhai)

Pseudocarchariidae

Pseudocarcharias

Crocodile shark (P. kamoharai)

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