Anthracotheriidae

Anthracotheriidae Fossil range: Middle Eocene–Late Pliocene PreЄ Є O S D C P T J K Pg N
Arsinoitherium (top) and Bothriogenys fraasi (bottom)
Scientific classification
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Artiodactyla Family: †Anthracotheriidae
Subfamilies and Genera[1][2]
†Anthracotheriinae †Anthracotherium (type genus) †Heptacodon †Siamotherium †Bothriodontinae †Aepinacondon †Afromeryx †Arretotherium †Brachyodus †Bothriogenys †Elomeryx †Hemimeryx †Kukusepasutanka †Libycosaurus †Merycopotamus †Sivameryx

Anthracotheriidae is a family of extinct, hippopotamus-like artiodactyl ungulates related to hippopotamuses and whales. The oldest genus, Elomeryx, first appeared during the Middle Eocene in Asia. They thrived in Africa and Eurasia, with a few species ultimately entering North America during the Oligocene, and eventually died out in Europe and Africa during the Miocene, possibly due to a combination of climatic changes and competition with other artiodactyls, including pigs and true hippopotami.^[3] The youngest genus, Merycopotamus, died out in Asia during the late Pliocene(change to "Pleistocene" ref- I have )). The family is named after the first genus discovered, Anthracotherium, which means "coal beast," as the first fossils of it were found in Paleogene-aged coal beds in France. Fossil remains of the anthracothere genus were discovered by the Harvard University and Geological Survey of Pakistan joint research project (Y-GSP) in the well-dated Middle and Late Miocene deposits of the Potwar Plateau in northern Pakistan.^[4]

In life, the average anthracothere would have resembled a skinny hippopotamus with a comparatively small, narrow head and most likely pig-like in general appearance.^[5] They had four or five toes on each foot, and broad feet suited to walking on soft mud. They had a full set of approximately 44 teeth (ref- I have)with five semi-crescentric cusps on the upper molars,^[3] which, in some species, were adapted for digging up the roots of aquatic plants.^[6]

Recent evidence gained from comparative gene sequencing further suggests that anthracotheres, as relatives of hippos,^[7] may be close to the ancestry of the whales.^[8][9] From the nature of the sediments in which they are fossilized, it seems that they were amphibious and this supports the view based on anatomical evidence that they were ancestors of the hippopotami.^[10] In many respects, especially the anatomy of the lower jaw, Anthracotherium, as with other members of the family, is allied to the hippopotamus, of which it is probably an ancestral form.^[11] However, the earliest known anthracotheres appear in the fossil record in the Middle Eocene, well after the archaeocetes had taken up a totally aquatic lifestyle. Although phylogenetic analyses of molecular data on extant animals strongly support the notion that hippopotamids are the closest relatives of cetaceans (whales, dolphins and porpoises) (REF), it is unlikely that they two groups are closely related when extant and extinct artiodactyls are analyzed. One reason hypothesizes that cetaceans originated about 50 million years ago in south Asia, whereas the family Hippopotimidae is only 15 million years old and the first hippopotamids are only 6 million years old. Yet, analyses of fossil clades have not resolved the issue of cetacean relations.^[12]

References

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