Capra

This article is about the Capra genus of animals and the Goat species. For other uses, see Capra, or Goat (disambiguation).

Capra
Female ibex
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Family:	Bovidae
Subfamily:	Caprinae
Genus:	Capra Linnaeus, 1758
Species
See Species and subspecies

Capra is a genus of mammals, the goats or wild goats, composed of up to nine species including the wild goat, the markhor and several species known as ibex.

The domestic goat (Capra aegagrus hircus) is a domesticated subspecies of the wild goat (Capra aegagrus).

Wild goats are animals of mountain habitats. They are very agile and hardy, able to climb on bare rock and survive on sparse vegetation.

The Rocky Mountain goat is in a separate genus, Oreamnos, and is not dealt with here.

Taxonomy

Range of the Capra-species

Male Alpine ibex

All members of the Capra genus are bovids (members of the family Bovidae) and caprids (subfamily Caprinae). All these are ruminants, meaning they chew the cud, and have a four-chambered stomach which plays a vital role in digesting, regurgitating and re-digesting their food.

The genus has sometimes been taken to include Ovis (sheep) and Ammotragus (Barbary Sheep),^[1] but these are usually regarded as distinct genera, leaving Capra for goats and ibexes. In this smaller genus, some authors have recognized only two species, the Markhor on one side and all other forms included in one species on the other side.^[2] Today nine species are usually accepted^[3]:

• Spanish ibex (Capra pyrenaica)
• Alpine ibex (Capra ibex)
• Nubian ibex (Capra nubiana)
• Siberian ibex (Capra sibirica)
• Walia ibex (Capra walie)
• West Caucasian tur (Capra caucasica)
• East Caucasian tur (Capra caucasica cylindricornis)
• Wild goat (Capra aegagrus)
  • Domestic goat (Capra aegagrus hircus sometimes refered to "Capra hircus")
• Markhor (Capra falconeri)

The goats of the genus Capra have complex systematic relationships, which are still not completely resolved. Recent studies based on mitochondrial DNA suggest that the Siberian ibex and the Nubian ibex represent distinct species, which are not very closely related to the physically similar Alpine ibex. The Alpine ibex forms a group with the Spanish ibex. The West Caucasian tur appears to be more closely related to the wild goat than to the East Caucasian tur. The markhor is relatively little separated from other forms – previously it had been considered to be a separate branch of the genus.^[4].

Almost all wild goat species are allopatric (geographically separated) – the only geographical overlaps are the wild goat (Capra aegagrus) with the East Caucasian tur (Capra cylindricornis), and the markhor (Capra falconeri) with the Siberian ibex (Capra siberica). In both cases the overlapping species do not usually interbreed in the wild – however, in captivity all Capra species can interbreed, producing fertile offspring.^[5]

Species and subspecies of goats

	Capra caucasica	West Caucasian tur
	Capra cylindricornis	East Caucasian tur
	Capra falconeri	Markhor
	Capra aegagrus	Wild goat
	Capra (aegagrus) hircus	domestic goat
	Capra sibirica	Siberian ibex
	Capra pyrenaica	Spanish ibex
	Capra walie	Walia ibex
	Capra ibex	Alpine ibex
	Capra nubiana	Nubian ibex

Domestication and uses

Main article: Domestic goat

Goats used for natural weed control

Along with sheep, goats were among the first domesticated animals. The domestication process started at least 10,000 years (calibrated calendar years) ago in what is now northern Iran^[6]. Easy human access to goat hair, meat, and milk were the primary motivations. Goat skins were popularly used until the Middle Ages for water and wine bottles when traveling and camping, and in certain regions as parchment for writing.

In ancient history

Evidence of the ibex is widely present in the archaeological record, particularly in the Near East and Mediterranean regions. Ibex motifs are very common on cylinder seals and pottery, both painted and embossed. Excavations from Minoan Crete at Knossos, for example, have yielded specimens from about 1800 BC, including one cylinder seal depicting an ibex defending himself from a hunting dog.^[7] From the similar age a gold jewelry ibex image was found at the Akrotiri archaeological site^[8] on Santorini in present day Greece.

An Iron Age Capra ibex specimen was recovered at the Aq Kupruk Archaeological site in present day Afghanistan, illustrating either domestication or hunting of the ibex by these early peoples.^{[clarification needed]} However, archaeological records of ibex can be difficult to separate from those of domestic goats.^[9]

Rock carvings of ibex as a charm to encourage childbearing or to give thanks for a child?

"Our Christian evangelist at Khalatse had become a father a few weeks before, and the people of the village had made presents of "flour-ibex" to him and his wife. He gave me one of those figures, which are made of flour and butter, and told me that it was a custom in Tibet and Ladakh, to make presents of "flour-ibex" on the occasion of the birth of a child. This is quite interesting information. I had often wondered why there were so many rock carvings of ibex at places connected with the pre-Buddhist religion of Ladakh. Now it appears probable that they are thank offerings after the birth of children. As I have tried to show in my previous article, people used to go to the pre-Buddhist places of worship, in particular, to pray to be blessed with children."^[10]

References

1. ^ Ansell, W. F. H. 1972. Order Artiodactyla. Part 15. Pp. 1-84, in The mammals of Africa: An identification manual (J. Meester and H. W. Setzer, eds.) [issued 2 May 1972]. Smithsonian Institution Press, Washington, D.C., not continuously paginated. (quoted in Grubb, Peter (16 November 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. ISBN 0-801-88221-4. http://www.bucknell.edu/msw3/browse.asp?id=14200767. )
2. ^ Haltenorth, T. 1963. Klassifikation der Säugetiere: Artiodactyla I. Handbuch der Zoologie, 8(32):1-167 (quoted in Grubb, Peter (16 November 2005). Wilson, D. E., and Reeder, D. M. (eds). ed. Mammal Species of the World (3rd edition ed.). Johns Hopkins University Press. ISBN 0-801-88221-4. http://www.bucknell.edu/msw3/browse.asp?id=14200767. )
3. ^ Nathalie Pidancier, Steve Jordan, Gordon Luikart, Pierre Taberlet: Evolutionary history of the genus Capra (Mammalia, Artiodactyla): Discordance between mitochondrial DNA and Y-chromosome. Molecular Phylogenetics and Evolution 40 (2006) 739–749 online
4. ^ Phylogenetic Reconstructions in the Genus Capra (Bovidae, Artiodactyla) Based on the Mitochondrial DNA Analysis. Russian Journal of Genetics, 2007, Vol. 43, No. 2, pp. 181–189. online
5. ^ V. G. Heptner: Mammals of the Sowjetunion Vol. I UNGULATES. Leiden, New York, 1989 ISBN 9004088741
6. ^ Melinda A. Zeder, Brian Hesse: The Initial Domestication of Goats (Capra hircus) in the Zagros Mountains 10,000 Years Ago. Science 24 March 2000: Vol. 287. no. 5461, pp. 2254 - 2257 online abstract
7. ^ C. Michael Hogan, Knossos fieldnotes, Modern Antiquarian (2007)
8. ^ M. Uda, G. Demortier, I. Nakai, X-rays for archaeology, 2005, Springer, 308 pages ISBN 1402035802
9. ^ Pam J. Crabtree, Douglas V. Campana, Kathleen Ryan, Early Animal Domestication and Its Cultural Context, 1989, University of Pennsylvania Museum of Archaeology ISBN 0924171960
10. ^ Francke, A. H. (1914). Antiquities of Indian Tibet. Two Volumes. Calcutta. 1972 reprint: S. Chand, New Delhi. pp. 95-96.

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