- In Irish mythology, the Milesians were the descendants of Míl Espáine, the final invaders of Ireland who defeated and displaced the semi-divine Tuatha Dé Danann.
- In the history of Hellenic civilization, the Milesians were the people of Miletus, a city in the Anatolia province of modern-day Turkey, who formed their own maritime empire throughout Turkey and into the Crimea.
- There are three philosophers from the aforementioned Hellenic civilization, Miletus, who are commonly referred to as the Milesian philosophers. They are Thales, Anaximander, and Anaximenes.
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