Scrophulariaceae

Figwort family
Scrophularia nodosa
Scientific classification
Kingdom: Plantae Division: Magnoliophyta Class: Magnoliopsida Order: Lamiales Family: Scrophulariaceae Juss.
Genera
See text.

Scrophulariaceae, the figwort family, is a family of flowering plants. The plants are annual or perennial herbs with zygomorphic (rarely actinomorphic) flowers. Members of the Scrophulariaceae have a cosmopolitan distribution, with the majority found in temperate areas, including tropical mountains. The family name is based on the name of the included genus Scrophularia L..

In the past it was treated as including about 275 genera and over 5,000 species, but its circumscription has been radically altered since numerous molecular phylogenies have shown the traditional broad circumscription to be grossly polyphyletic. Many genera have recently been transferred to other families within the Lamiales, notably Plantaginaceae and Orobanchaceae but also several new families (Olmstead et al., 2001; Olmstead, 2003). Several families of the Lamiales have had their circumscriptions enlarged to accommodate genera transferred from Scrophulariacae s.l.

The family includes some medicinal plants, among them:

• Leptandra, Black root, Culver's root, Leptandra
• Rehmannia, Chinese foxglove, used in Traditional Chinese medicine
• Scrophularia, figworts
• Verbascum, mulleins

Included genera

• Agalinis Raf. -- False foxglove
• Alectra Thunb.
• Alonsoa Ruiz & Pav.
• Aureolaria Raf. -- Yellow False foxglove
• Brachystigma Pennell -- Desert foxglove
• Buchnera L. -- Bluehearts
• Buddleja -- Butterfly-bush
• Capraria L.
• Chaenorrhinum -- Dwarf Snapdragon
• Dasistoma Raf.
• Diascia
• Diplacus Nutt. -- Bush monkeyflower
• Hemianthus Nutt.
• Ilysanthes
• Lendneria Minod
• Leptandra -- Black root, Culver's root, Leptandra)
• Leucospora Nutt.
  
• Limosella -> Plantaginaceae -> Scrophulariaceae (according to Oxelman et al. 2005)
• Macranthera Nutt.
• Mimetanthe Greene
• Nemesia
• Phygelius E.Mey. ex Benth. -- Cape fuchsia
• Rehmannia Libosch.
• Schwalbea L. -- Schwalbea
• Scrophularia L. -- Figwort
• Seymeria Pursh -- Black senna
• Sutera "Bacopa"
• Striga Lour. -- Witchweed
• Tomanthera
• Verbascum L. -- Mullein

Excluded genera

The following genera, traditionally included in the Scrophulariaceae, have been transferred to other families as indicated:

• Amphianthus -> Plantaginaceae
• Angelonia -> Plantaginaceae
• Antirrhinum -> Plantaginaceae
• Bacopa -> Plantaginaceae
• Bartsia -> Orobanchaceae
• Bellardia -> Orobanchaceae
• Besseya -> Plantaginaceae
• Calceolaria -> Calceolariaceae
• Castilleja -> Orobanchaceae
• Chaenorrhinum -> Plantaginaceae
• Chelone -> Plantaginaceae
• Chionophila -> Plantaginaceae
• Collinsia -> Plantaginaceae
• Cordylanthus -> Orobanchaceae
• Cymbalaria -> Plantaginaceae
• Digitalis -> Plantaginaceae
• Dopatrium -> Plantaginaceae
• Epixiphium -> Plantaginaceae
• Euphrasia -> Orobanchaceae
• Galvezia -> Plantaginaceae
• Gambelia -> Plantaginaceae
• Glossostigma -> Phrymaceae
• Gratiola -> Plantaginaceae
• Hebe -> Plantaginaceae
• Holmgrenanthe -> Plantaginaceae
• Howelliella -> Plantaginaceae
• Keckiella -> Plantaginaceae
• Kickxia -> Plantaginaceae
• Lagotis -> Plantaginaceae
• Leucophyllum -> Plantaginaceae
• Limnophila -> Plantaginaceae
• Linaria -> Plantaginaceae
• Lindernia -> Plantaginaceae, also placed within its own family Linderniaceae according to Haston et al 2007 (also known as LAPG II). ^[1]
• Lophospermum -> Plantaginaceae
• Mabrya -> Plantaginaceae
• Maurandella -> Plantaginaceae
• Maurandya -> Plantaginaceae
• Mazus -> Phrymaceae
• Mecardonia -> Plantaginaceae
• Melampyrum -> Orobanchaceae
• Micranthemum -> Plantaginaceae
• Mimulus -> Phrymaceae
• Misopates -> Plantaginaceae
• Mohavea -> Plantaginaceae
• Nemesia -> Plantaginaceae
• Neogaerrhinum -> Plantaginaceae
• Nothochelone -> Plantaginaceae
• Nuttallanthus -> Plantaginaceae
• Odontites -> Orobanchaceae
• Orobanche -> Orobanchaceae
• Orthocarpus -> Orobanchaceae
• Parentucellia -> Orobanchaceae
• Paulownia -> Paulowniaceae
• Pedicularis -> Orobanchaceae
• Penstemon -> Plantaginaceae
• Pseudorontium -> Plantaginaceae
• Rhinanthus -> Orobanchaceae
• Russelia -> Plantaginaceae
• Sairocarpus -> Plantaginaceae
• Schistophragma -> Plantaginaceae
• Scoparia -> Plantaginaceae
• Stemodia -> Plantaginaceae
• Synthyris -> Plantaginaceae
• Tonella -> Plantaginaceae
• Torenia L. -> Linderniaceae
• Triphysaria -> Orobanchaceae
• Veronica -> Plantaginaceae
• Veronicastrum -> Plantaginaceae

References

• Olmstead, R. G., dePamphilis, C. W., Wolfe, A. D., Young, N. D., Elisons, W. J. & Reeves P. A. (2001). "Disintegration of the Scrophulariaceae". American Journal of Botany 88: 348-361.  - on line here.
• Olmstead, R. G. (2003). "Whatever happened to the Scrophulariaceae?". Fremontia 30: 13-22.  - on line here.
• Oxelman, B.; Kornhall, P.; Olmstead, R.G.; Bremer, B. (2005). "Further disintegration of the Scrophulariaceae". Taxon 54: 411-425. 

External links

Wikimedia Commons has media related to:

Scrophulariaceae

• Kew list of Scrophulariaceae

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