An order of liverworts in the subclass Marchantiidae, consisting of two families, the terrestrial Sphaerocarpaceae (Sphaerocarpos and Geothallus) and the aquatic Riellaceae (Riella). The plants are characterized by envelopes surrounding each antheridium and archegonium, absence of elaters, poor development of seta, and absence of thickenings in the unilayered wall of an indehiscent capsule.
The rhizoids are smooth, and air chambers and pores are lacking. Oil bodies may be present. The antheridia are separately contained within receptacles on the dorsal surface or at the margins, while the archegonia are separately produced in receptacles dorsally or behind the growing point. The sporophyte consists of a bulbous foot, short or obsolescent seta, and globose indehiscent capsule with a unilayered wall. See also Bryophyta.