snout beetle

Weevil
Doydirhynchus austriacus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Cucujiformia
Superfamily:	Curculionidea Latreille, 1802
Families
Anthribidae — fungus weevils Attelabidae — leaf rolling weevils Belidae — primitive weevils Brentidae — straight snout weevils Caridae Curculionidae — true weevils Nemonychidae — pine flower weevils

A weevil is any beetle from the Curculionoidea superfamily. They are usually small, less than 6 mm (¼ inch), and herbivorous. There are over 60,000 species in several families, mostly in the family Curculionidae (the true weevils). Some other beetles, although not closely related, bear the name "weevil", such as the biscuit weevil (Stegobium paniceum), which belongs to the family Anobiidae.

Many weevils are damaging to crops. The grain or wheat weevil (Sitophilus granarius) damages stored grain. The boll weevil (Anthonomus grandis) attacks cotton crops. It lays its eggs inside cotton bolls, and the young weevils eat their way out.

Weevils are often found in dry foods including nuts and seeds, cereal and grain products, such as pancake mix. In the domestic setting, they are most likely to be observed when a bag of flour is opened. Their presence is often indicated by the granules of the infested item sticking together in strings, as if caught in a cobweb. If ingested, E. coli infection^{[citation needed]} and other various diseases^{[clarification needed]} can be contracted from weevils, depending on their diet.

Taxonomy

Because there are so many species and such diversity, the higher classification of weevils is in a state of flux. Weevils are generally divided into two major divisions, the Orthoceri or primitive weevils, and the Gonatoceri or true weevils (Curculionidae). E. C. Zimmerman proposed a third division, the Heteromorphi, for several intermediate forms.^[1] Primitive weevils are distinguished by having straight antennae, while true weevils have elbowed (geniculate) antennae. The elbow occurs at the end of the scape (first antennal segment) in true weevils, and the scape is usually much longer than the other antennal segments. Some exceptions occur. Nanophyini are primitive weevils (with very long trochanters) but have long scapes and geniculate antennae. From the true weevils, Gonipterinae and Ramphus have short scapes and little or no elbow.

The most recent classification system to family level was provided by Kuschel,^[2] with updates from Marvaldi et al.,^[3] and was achieved using phylogenetic analyses. The accepted families are the primitive weevils, Anthribidae, Attelabidae, Belidae, Brentidae, Caridae and Nemonychidae, and the true weevils Curculionidae. Most other weevil families were demoted to subfamilies or tribes. Weevil species radiation was shown to follow steps in plant evolution upon which the weevils feed.

Some of the features used to distinguish weevil families are:

Phylogeny

A Long Nosed Weevil, Rhinotia hemistictus from the family Belidae

A phylogeny of the Curculionidae based on 18S ribosomal DNA and morphological data is suggested below:^[3]

References

• Bright, Donald, E., Bouchard, P. (2008) Coleoptera, Curculionidae, Entiminae. Weevils of Canada and Alaska — Volume 2. Insects and Arachnids of Canada Series, Part 25 Ottawa: NRC Research Press. ISBN 978-0-660-19400-4.

Gallery

Stout beetle (Lixus angustatus)	Acorn weevil, Curculio sp.	A weevil of the Belidae family on parsley flowers	A weevil from India
An attelabid weevil from India	Palmetto weevil	Clover leaf weevil

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