Grapevine is an informal network of communication within an organisation, also termed by many as"rumors" or "gossip", such as over the water-cooler communication etc.
Around 70% of all communication in an organisation is done through the grapevine network and it is considered by researchers to be highly accurate.
The Grapevine is an informal communication network that moves throughout an organization. It develops when humans interact. It's the rumor connection. When employees feel left-out, they take cover in the Grapevine. It's the opinion place when some employees are too afraid to confront the formal channel of communication.
I'm assuming here that you do not mean the living plant that grapes grow on, since you are in the language category; in which case "Grapevine" refers to, usually means, a network of gossip, word-of-mouth discussion about a particluar subject. Prime example is the title and words of the American song, "I heard it through the Grapevine" by Marvin Gaye, originally, and later by other fine musical artists. Another example: I heard it through the grapevine that there's going to be a pop quiz in class today. Derivation: "Powderpuff (women's makeup tool) Grapevine" Refers to a network, usually gossip consisting of females exclusively: Powderpuff Grapevine. This could be considered misogynistic, however, so be cautious when using this or any other form of gossip.
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