Traditionally, it is the very top tier or the small piece of cake that help the cake topper that the bride and groom keep for memorie's sake.
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It's for the bride and groom to eat themselves.
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A tier of a cake is a layer.
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A tier is a section on a cake, so a mid-tier would be the middle section on a cake.
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The Christians' wedding cake is one big affair which originated primarily with lots of little wheat cakes. These little cakes were broken over the bride's head to bring good luck and fertility. The present guests then eat a crumb to make it ensure good luck. The vogue of three tier wedding cake started off after the bizarre shape of the spire of Saint Bride's Church. The cutting of the cake together implies that the "just married couple" would share their life together. Tradition has it that the top layer represents them as a couple, while the bottom layer represents the couple as a family. The layer in between represent the number of children the couple would have.
There are not too many rules to follow. Usually, take a slice (reasonable size) out of the bottom tier or tier just below arm level. The bride and groom get the first slice, if they are not the ones cutting the cake themselves. It would also be nice, if you are cutting the cake for them, to wish the bride and the groom a healthy and happy marriage.
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