What is a epsilon in a wolf pack?

Answer 1

Gamma is not an official title in ranking wolves.

Alpha is the pack leader, or pack leading pair (male and female) and typically the only two mating in the pack. (the rest often being children of the alpha's)

The beta wolf is second in command, the strongest next to the alpha and the one who would be most likely to take charge if the alpha died.

The omega wolf is the weakest of the pack, or the runt in a litter. Typically the weakest and least cared for the omega may not even be fed if there is a limited food supply and not enough to go around.

Beyond that any labels like gamma would be up to the individual observer to use, some wolf researchers may use the entire Greek alphabet (alpha, beta, gamma, delta, etc) simply to label every member in a pack but this is up to the individual researcher if they choose to give names, or attempt to recognize where every individual within the pack stands.

There is typically only the alpha pair, the beta second and the omega runt which are clear to observation; the rest of the pack falling somewhere in between with relatively equal status. You could in theory call all of those in between the beta and the omega: 'the gamma wolves'; but like I said, gamma hasn't really caught on as a regular part of the lexicon of wolf observation.