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Evolution has two key driving forces without which it would not occur, but with which it must occur:

The first is reproductive variation: this basically asserts that each organism is slightly different from both its parents and its siblings.

The second is differential reproductive success: this asserts that features that allow their bearers to reproduce more successfully than members of the same species with different features are likely to be present in a greater part of the population in coming generations.

In other words: the key driving forces behind evolution are variation and selection.

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