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The quote was coined by Saint Ambrose (340AD - 397AD), the Archbishop of Milan. The full text was: "si fueris Romae, Romano vivito more; si fueris alibi, vivito sicut ibi" - which translated to English means: "if you are in Rome, live in the Roman way; if you are elsewhere, live as they do there".
As time went on, people twisted the original translation to "When in Rome, do as the Romans do". But the original version was as above.
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