A440, also known as the A above "middle C", and further known as
the note most commonly associated with 440 hertz was made the
international standard for pitch decided upon in 1955 (ISO 16), and
was reaffirmed by the same organization in 1975.
http://en.wikipedia.org/wiki/Pitch_(music)
I can't find any reasoning as to why A has always traditionally
been used for the pitch standard. Even when the tuning fork was
originally invented, an A was chosen as the best tuning note. My
best guess is that it is due to the note's alphabetic
significance.
It should also be noted that although some orchestras tune to an
electronic tone that is calibrated to 440 hertz, many do not. Many
orchestras are primarily concerned with whether or not they're in
tune with each other, and as such, the entire orchestra (or band)
often tune to an oboe. The oboe cannot change its tuning, so
oftentimes the other members of the band tune to it.
One of the primary reasons for an orchestra, or any group for
that matter, to tune to A440 is when there is an instrument whose
tuning cannot easily be changed, such as a piano, or organ with a
fixed tuning to A440.