Articulata
(invertebrate zoology) A class of the Brachiopoda having hinged valves that usually bear teeth. The only surviving subclass of the echinoderm class Crinoidea.
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(invertebrate zoology) A class of the Brachiopoda having hinged valves that usually bear teeth. The only surviving subclass of the echinoderm class Crinoidea.
The only surviving subclass of the Crinoidea. It differentiated during Triassic times. The calyx is dicyclic, but considerable reduction of the infrabasals and basals may occur. The uniserial arms bear pinnules and usually branch, and the arm retains its movable articulation with the radial plate, despite the incorporation of the lower brachial ossicles into the calyx. Extant stalked forms with nodal rings of cirri (Metacrinus) are included in the order Isocrinida (see illustration). They do not tolerate turbulent waters and live at depths below current action, although they do inhabit shallow water when the conditions are suitable. The feather stars, of the order Comatulida, discard the stem when young, and thereafter remain free, either as swimming animals or as creeping benthic forms. They prefer shallow, clear water, rich in nutrients, and therefore abound on tropical coasts and in polar seas rather than in temperate waters. Four other orders have been defined. See also Crinoidea; Echinodermata.
articulate stalked crinoid.">
Metracrinus cyaneus, a living, articulate stalked crinoid.
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A Passion Flower feather star
(Ptilometra australis) |
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Bourgeticrinida |
Articulata are the only extant subclass of the class Crinoidea. They are differentiated
from the extinct subclasses by their lack of an anal plate and the presence of entoneural system. Articulata are stalked
echinoderms with pentamerous symmetry. Their stalk, which consists of numerous disks held together by ligaments, supports an
aboral cup made of circlets of calcerous plates. five arms, which consist of articulated series of ossicles, extend from the
aboral plate and form the food-capture mechanism of articulata. The arms of articulata are pinnulate in that they have
alternating pinnules branching out along them to effectively increase the surface area for feeding. These pinnules all have
ciliated ambulacral grooves that converge to form larger grooves in the arms that lead down to the mouth located beside the anus
on the upper surface of the aboral plate. Articulata are passive suspension feeders. They capture algea with triplets of tube
feet located on the pinnules, and the ciliated ambulacral canals transport this algae to the mouth. Although they are passive
feeders, some articulata have been observed to move to better feeding areas either with locomotory mechanisms at the base of the
stalk or by detaching and pulling themselves with their arms. Currently there are 540 described species of articulata that fall
into two major groups. The bourgueticrinids which have the traditional stalked body form account for about 15% percent of the
known species.
Articulata first appeared in the fossil record during the Triassic period although other, now extinct crinoid groups, originated in the Ordivician.
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