seagull

Gulls
Black-headed Gull
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Charadriiformes
Suborder:	Lari
Family:	Laridae Vigors, 1825
Genera
Larus Ichthyaetus Chroicocephalus Saundersilarus Leucophaeus Hydrocoloeus Rissa Pagophila Rhodostethia Xema Creagrus

Gulls (often informally called seagulls) are birds in the family Laridae. They are most closely related to the terns (family Sternidae) and only distantly related to auks, skimmers, and more distantly to the waders. Until recently, most gulls were placed in the genus Larus, but this arrangement is now known to be polyphyletic, leading to the resurrection of several genera.^[1]

They are typically medium to large birds, usually grey or white, often with black markings on the head or wings. They typically have harsh wailing or squawking calls. They have stout, longish bills, and webbed feet. Gull species range in size from the Little Gull, at 120 g (4.2 oz) and 29 cm (11.5 inches), to the Great Black-backed Gull, at 1.75 kg (3.8 lbs) and 76 cm (30 inches).

Biology and habits

Most gulls, particularly Larus species, are ground nesting carnivores, which will take live food or scavenge opportunistically. Live food often includes crabs and small fish. Apart from the kittiwakes, gulls are typically coastal or inland species, rarely venturing far out to sea.

The large species take up to four years to attain full adult plumage, but two years is typical for small gulls. Large White-Headed Gulls are typically long-lived birds, with a maximum age of 49 years recorded for the Herring Gull.^[2]

Gulls nest in large, densely packed, and noisy colonies. They lay two to three speckled eggs in nests composed of vegetation. The young are precocial, being born with dark mottled down, and mobile from birth^[3].

Gulls—the larger species in particular—are resourceful, inquisitive and highly intelligent birds, demonstrating complex methods of communication and a highly developed social structure; for example, many gull colonies display mobbing behaviour, attacking and harassing would-be predators and other intruders.^[4] In addition, certain species (e.g. the Herring Gull) have exhibited tool use behaviour. Many species of gull have learned to coexist successfully with humans and have thrived in human habitats.^[5] Others rely on kleptoparasitism to get their food. Gulls have been observed preying on live whales, landing on the whale as it surfaces to peck meat,^[6]

Taxonomy

The taxonomy of gulls is confused by their widespread distribution and geneflow leading to zones of hybridization. Some have traditionally been considered ring species, but recent evidence suggest this assumption is questionable.^[7] Until recently, most gulls were placed in the genus Larus, but this arrangement is now known to be polyphyletic, leading to the resurrection of the genera Ichthyaetus, Chroicocephalus, Leucophaeus, Saundersilarus and Hydrocoloeus.^[1] Some English names refer to species complexes within the group:

• Large white-headed gull is used to describe the 18 or so Herring Gull-like species from California Gull to Lesser Black-backed Gull in the taxonomic list below.
• White-winged gull is used to describe the 4 pale-winged, High Arctic-breeding taxa within the former group; these are Iceland Gull, Glaucous Gull, Thayer's Gull, and Kumlien's Gull.

Hybridisation between species of gull occurs quite frequently, although to varying degrees depending on the species involved (see Hybridisation in gulls). The taxonomy of the large white-headed gulls is particularly complicated.

In common usage, members of various gull species are often referred to as sea gulls or seagulls. This name is used by the layman to refer to a common local species or all gulls in general, and has no fixed taxonomic meaning.

The American Ornithologists' Union combines Sternidae, Stercorariidae, and Rhynchopidae as subfamilies in the family Laridae, but recent research ^[8][9][10] indicates that this is incorrect.

List of species

This is a list of gull species, presented in taxonomic sequence.

Two gulls fighting over a freshly-killed ground squirrel

A gull landing on a sea otter, probably trying to steal its prey

Gulls also steal food from people

A gull eating a starfish

A gull eating a crayfish

A gull attacking a coot

Gulls in a feeding frenzy

Gulls feeding on Elephant Seal placenta and membranes

Seagull dining on remains of eels left in fishing nets

Genus Larus

• Pacific Gull, Larus pacificus
• Belcher's Gull, Larus belcheri
• Olrog's Gull, Larus atlanticus
• Black-tailed Gull, Larus crassirostris
• Heermann's Gull, Larus heermanni
• Common Gull or Mew Gull, Larus canus
• Ring-billed Gull, Larus delawarensis
• California Gull, Larus californicus
• Great Black-backed Gull, Larus marinus
• Kelp Gull, Larus dominicanus (called "Southern Black-backed Gull" or "Karoro" in New Zealand)
  • Cape Gull, Larus dominicanus vetula
• Glaucous-winged Gull, Larus glaucescens
• Western Gull, Larus occidentalis
• Yellow-footed Gull, Larus livens
• Glaucous Gull, Larus hyperboreus
• Iceland Gull, Larus glaucoides
  • Kumlien's Gull, Larus glaucoides kumlieni
• Thayer's Gull, Larus thayeri
• Herring Gull, Larus argentatus
• Heuglin's Gull, Larus heuglini
• American Herring Gull, Larus smithsonianus
• Yellow-legged Gull, Larus michahellis
• Caspian Gull, Larus cachinnans
• East Siberian Herring Gull, Larus vegae
• Armenian Gull, Larus armenicus
• Slaty-backed Gull, Larus schistisagus
• Lesser Black-backed Gull, Larus fuscus

Genus Ichthyaetus

• White-eyed Gull, Ichthyaetus leucophthalmus
• Sooty Gull, Ichthyaetus hemprichii
• Great Black-headed Gull, Ichthyaetus ichthyaetus
• Audouin's Gull, Ichthyaetus audouinii
• Mediterranean Gull, Ichthyaetus melanocephalus
• Relict Gull, Ichthyaetus relictus

Genus Leucophaeus

• Dolphin Gull, Leucophaeus scoresbii
• Laughing Gull, Leucophaeus atricilla
• Franklin's Gull, Leucophaeus pipixcan
• Lava Gull, Leucophaeus fuliginosus
• Gray Gull, Leucophaeus modestus

Genus Chroicocephalus

• Silver Gull, Chroicocephalus novaehollandiae
• Red-billed Gull, Chroicocephalus scopulinus
• Hartlaub's Gull, Chroicocephalus hartlaubii
• Brown-hooded Gull, Chroicocephalus maculipennis
• Gray-hooded Gull, Chroicocephalus cirrocephalus
• Andean Gull, Chroicocephalus serranus
• Black-billed Gull, Chroicocephalus bulleri
• Brown-headed Gull, Chroicocephalus brunnicephalus
• Black-headed Gull, Chroicocephalus ridibundus
• Slender-billed Gull, Chroicocephalus genei
• Bonaparte's Gull, Chroicocephalus philadelphia

Genus Saundersilarus

• Saunders's Gull, Saundersilarus saundersi

Genus Hydrocoloeus

• Little Gull, Hydrocoloeus minutus

Genus Rhodostethia

• Ross's Gull, Rhodostethia rosea

Genus Rissa

• Black-legged Kittiwake, Rissa tridactyla
• Red-legged Kittiwake, Rissa brevirostris

Genus Pagophila

• Ivory Gull, Pagophila eburnea

Genus Xema

• Sabine's Gull, Xema sabini

Genus Creagrus

• Swallow-tailed Gull, Creagrus furcatus

Evolution

The Laridae are known from fossil evidence since the Early Oligocene, some 30-33 mya. A fossil gull from the Middle to Late Miocene of Cherry County, Nebraska, USA is placed in the prehistoric genus Gaviota; apart from this and the undescribed Early Oligocene fossil, all prehistoric species were tentatively assigned to the modern genus Larus. Among those of them that have been confirmed as gulls, "Larus" elegans and "L." totanoides from the Late Oligocene/Early Miocene of southeast France have since been separated in Laricola.

References

Bibliography

• Grant, Peter J. (1986) Gulls: a guide to identification ISBN 0-85661-044-5
• Howell, Steve N. G. and Jon Dunn (2007) Gulls of the Americas ISBN 0-618-72641-1
• Olsen, Klaus Malling & Larsson, Hans (1995): Terns of Europe and North America. Christopher Helm, London. ISBN 0-7136-4056-1

External links

Look up gull in Wiktionary, the free dictionary.

Wikimedia Commons has media related to: Laridae

• Rudy's Gull-index Pictures of less well-known plumages of large gulls
• Gull videos on the Internet Bird Collection
• Gullpix Pictures of gulls, sorted by species