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For more information on Luigi Galvani, visit Britannica.com.
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[b. Bologna, (Italy) September 9, 1737, d. Bologna, December 4, 1798]
While studying movement in frogs, Galvani discovered that muscles of a frog's severed leg contract when in contact with two dissimilar metals because electric current is produced. Galvani incorrectly believed that the frog's tissues generated the electricity and the metals discharged it. In 1800 Alessandro Volta showed that electricity is produced from the contact of two different metals in a moist environment. By analogy with the frog's twitch, a person is said to be galvanized by any stimulus that causes a strong reaction.
| Biography: Luigi Galvani |
The Italian physiologist Luigi Galvani (1737-1798) is noted for his discovery of animal electricity.
Luigi Galvani was born at Bologna on Sept. 9, 1737. He studied theology for a while and then medicine at the University of Bologna. In 1762, upon completion of his studies, he was appointed lecturer of anatomy and surgery at Bologna. His interest focused on the animal senses, which led him into deep theoretical interest in the action of the nervous system.
By the middle of the 18th century various books on electricity were available in Italian, and in 1744 Benjamin Franklin's famous book on electricity appeared in Italian translation. Galvani was influenced by Franklin's "one fluid theory," according to which electrical phenomena were caused by an electric fluid that results in so-called positive electricity, while so-called negative electricity was the absence of fluid. What seemed especially important to Galvani was Franklin's explanation of the Leyden jar, the early form of condenser. According to Franklin, positive electricity accumulated on the inner conductor while the outer conductor became negatively charged. The whole setup was similar to an accumulation of fluid on the inside of the bottle. Galvani drew an analogy between the Leyden jar and animal muscle and carried out his experiments with this thought in mind. He studied the effects of electricity from lightning on muscular contractions in a frog and proved that the electricity produced muscular convulsions.
Galvani's first announcement of his experiments appeared in a paper, "On the Effect of Electricity on Muscular Motion," published in 1791. He also gave an account of convulsions produced in a frog, in the absence of an electrical machine, when the frog formed part of a circuit containing one or more pieces of metal. Galvani had observed motion of the nerve juices during these convulsions and proposed the theory that the convulsions were caused by electricity within the animal's body; the muscle fiber and the nerve were acting like a Leyden jar.
Galvani's great Italian contemporary Alessandro Volta began working on animal electricity in 1792 and came out in direct opposition to Galvani's theory of an animal electrical fluid. It was then that the famous controversy between the two began. Volta proved that the nerves were nothing but electrical conductors and that it was possible to get electrical effects by placing any two metals in contact with an intervening piece of moistened cardboard. In this controversy Volta was correct in his physical interpretations, yet it was Galvani's influence which fostered the flourishing science of neurophysiology in the 19th century. However, the controversy between the two men spread into their personal relations and even into Italian politics of the time.
After the Cisalpine Republic was established in 1797, Galvani refused to swear allegiance to it on religious grounds and was dismissed from his university position. Volta swore allegiance and played a central role in the republic. Galvani was reinstated a year later, but by then he was a completely broken man. He died on Dec. 4, 1798.
Further Reading
Biographical material on Galvani is in Bern Dibner, Galvani-Volta: A Controversy That Led to the Discovery of Useful Electricity (1952). See also James R. Partington, A History of Chemistry, vol. 4 (1964).
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| Luigi Galvani | |
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Luigi Galvani - Italian physician famous for pioneering bioelectricity.
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| Born | September 9, 1737 Bologna, Papal States |
| Died | December 4, 1798 (aged 61) Bologna, Papal States |
| Institutions | University of Bologna |
| Known for | Bioelectricity |
Luigi Galvani (September 9, 1737 – December 4, 1798) was an Italian physician and physicist who lived and died in Bologna. In 1771, he discovered that the muscles of dead frogs twitched when struck by a spark.[1] This was one of the first forays into the study of bioelectricity, a field that still today studies the electrical patterns and signals of the nervous system.
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At first he thought about being a mechanic because he loved taking and dealing with spool testing. Later, his wish was to enter the church, but by his parents he was educated for a medical career. Galvani attended Bologna's medicine school and became a medical doctor like his father. At the University of Bologna, he was in 1762 appointed public lecturer in anatomy, and soon gained repute as a skilled though not eloquent teacher, and, chiefly from his researches on the organs of hearing and genito-urinary tract of birds, as a comparative anatomist.
His celebrated theory of animal electricity he enunciated in a treatise, De viribus electricitatis in motu musculari commentarius published in the 7th volume of the memoirs of the Institute of Sciences at Bologna in 1791, and separately at Modena in the following year, and elsewhere subsequently. In 1764, he married Lucia Galleazzi, a daughter of a professor at the University of Bologna and a well-liked woman of society. In 1772 Galvani became president of the University.
According to popular version of the story, Galvani dissected a frog at a table where he had been conducting experiments with static electricity. Galvani's assistant touched an exposed sciatic nerve of the frog with a metal scalpel, which had picked up a charge. At that moment, they saw sparks and the dead frog's leg kick as if in life. The observation made Galvani the first investigator to appreciate the relationship between electricity and animation — or life. This finding provided the basis for the current understanding that electrical energy (carried by ions), and not air or fluid as in earlier balloonist theories, is the impetus behind muscle movement. He is poorly credited with the discovery of bioelectricity.
Galvani called the term animal electricity to describe whatever it was that activated the muscles of his specimens. Along with contemporaries, he regarded their activation as being generated by an electrical fluid that is carried to the muscles by the nerves. The phenomenon was dubbed galvanism, after Galvani, on the suggestion of his peer and sometime intellectual adversary Alessandro Volta. Galvanism is electricity by a medical reaction.
Galvani's investigations led shortly to the invention of an early battery, but not by Galvani, who did not perceive electricity as separable from biology. Galvani did not see electricity as the essence of life, which he regarded vitalistically. Galvani believed that the animal electricity came from the muscle. Galvani's associate Alessandro Volta, in opposition, reasoned that the animal electricity was a physical phenomenon, a metallic electricity.
While, as Galvani believed, all life is indeed electrical, specifically that all living things are made of cells and every cell has a cell potential, biological electricity has the same chemical underpinnings as the current between electrochemical cells, and thus can be recapitulated in a way outside the body, Volta's intuition was correct. Volta, essentially, objected to Galvani’s conclusions about "animal electric fluid", but the two scientists disagreed respectfully and Volta coined the term "galvanism" for a direct current of electricity produced by chemical action.[2] Thus, owing to an argument between the two in regard to the source or cause of the electricity, Volta built the first battery in order to specifically disprove his associate's theory. Volta's “pile” became known therefore as a voltaic pile.
Galvani’s home in Bologna has been preserved and can be seen in the central via Marconi. On the facade of the house, now a seat of a bank, there is a medallion with the face of Galvani and double inscription in Italian and Latin: "NATO ACCOLSI GALVANI E PIANSI ESTINTO. PER LUI FU L'UNO ALL'ALTRO POLO AVVINTO - GALVANUM EXCEPI NATUM LUXIQUE PEREMPTUM CUIUS AB INVENTO IUNCTUS UTERQUE POLUS" (I received the newborn Galvani; I cried him dead / He held together both the electric poles).
Galvani’s monument. In the square dedicated to him, facing the palace of the Archiginnasio, the ancient seat of the University of Bologna, a big marble statue has been erected to the scientist while observing one of his famous frog preparation.
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