pine¹

 

common name for members of the Pinaceae, a family of resinous woody trees with needlelike, usually evergreen leaves. The Pinaceae reproduce by means of cones (see cone) rather than flowers and many have winged seeds, suitable for wind distribution. They are found chiefly in north temperate regions, where they form vast forests. The family was apparently more abundant in the mid-Cenozoic era, but it has maintained its population better than other gymnosperms because the trees are more adaptable to cold, dry climates; the reduced leaf surface and deep-set stomata minimize loss of water by transpiration. The family is the largest and most important of the conifers, providing naval stores, paper pulp, and more lumber by far than any other family. In some localities almost pure stands occur, permitting economical lumbering of large numbers of a given type of tree. Of the family's nine genera four are widely dispersed throughout North America and the Old World. Members of all nine genera are represented in horticulture as introduced timber trees or ornamentals. The so-called kauri pine, although pinelike in appearance, belongs to another family (see monkey-puzzle tree).

The True Pines

Pinus (the true pines) is the largest and most widespread genus, characteristic of many north temperate regions (except the plains), especially at lower altitudes, and in a few tropical regions, notably on mountain slopes. Species of Pinus can often be identified by the leaf arrangement, one needle or clusters of from two to five (in all cases enclosed in a sheath at the base) being consistently produced by each type. Many of the pines are economically valuable; from them come the naval stores: pitch (see tar and pitch), turpentine, and rosin. Drying and nondrying oils are also made from the seeds of some pines. Several Mediterranean and American species yield edible seeds (see pine nut).

The ponderosa pine or western yellow pine (P. ponderosa), is a hard pine second only to the Douglas fir as a commercial timber tree in North America. The white pine (P. strobus) has straight-grained soft wood with little resin, used especially for interior trim and cabinetwork. It once grew densely from Newfoundland to Manitoba and over much of the E United States westward to Minnesota, but constant felling and attacks of white-pine blister rust have greatly depleted the stands, especially in the NE United States. The Norway pine, or red pine, (P. resinosa) has a similar range and has also suffered from overcutting. Its wood is somewhat heavier and is suitable for general construction. The Norway pine is frequently used in reforestation programs. The jack pine (P. banksiana), the most northern of the American species, thrives on poor and sandy soils and is much used to colonize areas where more valuable species may later be introduced. Although the trunk is often gnarled, making it unsuitable for good lumber, it supplies much pulpwood and is used locally for rough lumber, fuel, and crating. The Virginia pine (P. virginiana) of the Appalachians and the Piedmont is popular regionally as a Christmas tree. The longleaf pine, or Southern yellow pine (P. palustris) has highly resinous wood used for heavy construction and as a major source of naval stores and pulpwood. It and the faster growing slash pine (P. caribaea) of the same region have gained importance as northern pine stands have been depleted. The latter is widely cultivated in tropical areas with sandy soils. The Scotch pine (P. sylvestris), ranging from Scotland to Siberia and popular as a Christmas tree in the United States, is one of the most valuable timber trees of Europe. The cluster pine (P. pinaster), widespread in S France and in Spain, is the chief European source of turpentine. The Monterey pine (P. radiata) of California has been widely planted in New Zealand and Chile for reforestation.

Other Species in the Pine Family

Abies (fir) species are usually of more northern distribution and found at higher altitudes. Sap-filled “blisters” on the trunks of some species provide balsam. Larix (larch) and Pseudolarix (golden larch, of China) are the only two deciduous genera. Picea (spruce) is the world's most important source of paper. Cedrus (cedar) ranges from the Mediterranean area to the Himalayas; Keteleeria is restricted to E and SE Asia.

Tsuga (hemlock) and Pseudotsuga are native only to North America and E Asia. Pseudotsuga menziesii (the Douglas fir) of W North America, one of the tallest trees known (up to 385 ft/117 m) and the leading timber-producing tree of the continent, is carefully controlled by forestry measures. Its wood, usually hard and strong, is of great commercial importance for construction; it is also commonly used as a Christmas tree in the United States. Named for David Douglas, the tree has many local names, e.g., Douglas spruce, Oregon pine, red fir, and yellow fir.

Classification

Pines are classified in the division Pinophyta, class Pinopsida, order Coniferales.

Pine
Maritime Pine (Pinus pinaster)
Scientific classification
Kingdom: Plantae Division: Pinophyta Class: Pinopsida Order: Pinales Family: Pinaceae Genus: Pinus L.
Subgenera
Subgenus Strobus Subgenus Ducampopinus Subgenus Pinus See Pinus classification for complete taxonomy to species level. See list of pines by region for list of species by geographical distribution.

A pine is a coniferous tree in the genus Pinus, in the family Pinaceae. They make up the monotypic subfamily Pinoideae. There are about 115 species of pine, although different authors accept anything from 105 to 125 species.

Distribution

Pines are native to most of the Northern Hemisphere. In Eurasia, they range from the Canary Islands and Scotland east to the Russian Far East, and the Philippines, north to just over 70°N in Norway (Scots Pine) and eastern Siberia (Siberian Dwarf Pine), and south to northernmost Africa, the Himalaya and Southeast Asia, with one species (Sumatran Pine) just crossing the Equator in Sumatra to 2°S. In North America, they range from 66°N in Canada (Jack Pine) south to 12°N in Nicaragua (Caribbean Pine). The highest diversity in the genus occurs in Mexico and California.

Pines have been introduced in subtropical and temperate portions of the Southern Hemisphere, including Chile, Brazil, Australia, and New Zealand, where they are grown widely as a source of timber, and some are becoming invasive species.

Morphology

Pines are evergreen and resinous trees (rarely shrubs) growing to 3–80 m tall, with the majority of species reaching between 15-45 m tall. The smallest are Siberian Dwarf Pine and Potosi Pinyon, and the tallest, Sugar Pine. Pines are long-lived, typically reaching ages of 100–1,000 years, and some even longer; the longest-lived is Great Basin Bristlecone Pine, where one individual at 4,839 years (2007) is the oldest living organism in the world.

The bark of most pines is thick and scaly, but some species have thin, flaking bark. The branches are produced in regular "pseudowhorls", actually a very tight spiral but appearing like a ring of branches arising from the same point. Many pines are uninodal, producing just one such whorl of branches each year, from buds at the tip of the year's new shoot, but others are multinodal, producing two or more whorls of branches per year. The spiral growth of branches, needles and cone scales are arranged in Fibonacci number ratios. The new spring shoots are sometimes called "candles"; they are covered in brown or whitish bud scales and point upward at first, then later turn green and spread outward. These "candles" offer foresters a means to evaluate fertility of the soil and vigour of the trees.

Foliage

Pines have four types of leaves:

1. Seed leaves (cotyledons) on seedlings, borne in a whorl of 4-24.
2. Juvenile leaves, which follow immediately on seedlings and young plants, 2-6 cm long, single, green or often blue-green, and arranged spirally on the shoot. These are produced for six months to five years, rarely longer (and also produced later in life after injury in some pines).
3. Scale leaves, similar to bud scales, small, brown and non-photosynthetic, and arranged spirally like the juvenile leaves.
4. Needles, the adult leaves, which are green (photosynthetic), bundled in clusters (fascicles) of (1-) 2-5 (-6) needles together, each fascicle produced from a small bud on a dwarf shoot in the axil of a scale leaf. These bud scales often remain on the fascicle as a basal sheath. The needles persist for 1.5-40 years, depending on species. If a shoot is damaged (e.g. eaten by an animal), the needle fascicles just below the damage will generate a bud which can then replace the lost growth.

Cones

Pines are mostly monoecious, having the male and female cones on the same tree, though a few species are sub-dioecious with individuals predominantly, but not wholly, single-sex. The male cones are small, typically 1-5 cm long, and only present for a short period (usually in spring, though autumn in a few pines), falling as soon as they have shed their pollen. The female cones take 1.5-3 years (depending on species) to mature after pollination, with actual fertilization delayed one year. At maturity the cones are 3-60 cm long. Each cone has numerous spirally arranged scales, with two seeds on each fertile scale; the scales at the base and tip of the cone are small and sterile, without seeds. The seeds are mostly small and winged, and are anemophilous (wind-dispersed), but some are larger and have only a vestigial wing, and are bird-dispersed (see below). At maturity, the cones usually open to release the seeds, but in some of the bird-dispersed species (e.g. Whitebark Pine), the seeds are only released by the bird breaking the cones open. In others, the fire climax pines (e.g. Monterey Pine, Pond Pine), the seeds are stored in closed ("serotinous") cones for many years until a forest fire kills the parent tree; the cones are also opened by the heat and the stored seeds are then released in huge numbers to re-populate the burnt ground.

Classification

Pines are divided into three subgenera, based on cone, seed and leaf characters:

• Subgenus Strobus (white or soft pines). Cone scale without a sealing band. Umbo terminal. Seedwings adnate. One fibrovascular bundle per leaf.
• Subgenus Ducampopinus (pinyon, lacebark and bristlecone pines). Cone scale without a sealing band. Umbo dorsal. Seedwings articulate. One fibrovascular bundle per leaf.
• Subgenus Pinus (yellow or hard pines). Cone scale with a sealing band. Umbo dorsal. Seedwings articulate. Two fibrovascular bundles per leaf.

Ecology

Pines grow well in acid soils, some also on calcareous soils; most require good soil drainage, preferring sandy soils, but a few, e.g. Lodgepole Pine, will tolerate poorly drained wet soils. A few are able to sprout after forest fires, e.g. Canary Island Pine. Some species of pines, e.g. Bishop Pine, need fire to regenerate and their populations slowly decline under fire suppression regimes. Several species are adapted to extreme conditions imposed by elevation and latitude; see e.g. Siberian Dwarf Pine, Mountain Pine, Whitebark Pine and the bristlecone pines. The pinyon pines and a number of others, notably Turkish Pine, are particularly well adapted to growth in hot, dry semi-desert climates.

The seeds are commonly eaten by birds and squirrels. Some birds, notably the Spotted Nutcracker, Clark's Nutcracker and Pinyon Jay, are of importance in distributing pine seeds to new areas where they can grow. Pine needles are sometimes eaten by some Lepidoptera species (see list of Lepidoptera which feed on Pines) and also the Symphytan species Pine Sawfly.

Uses

Pines are commercially among the most important of species used for timber and wood pulp in temperate and tropical regions of the world. This is because they are fast-growing softwoods that can be planted in relatively dense stands, and because their acidic decaying needles may inhibit the growth of other competing plants in the cropping areas. Commercial pine crops are grown in plantations.

The resin of some species is important as the source of turpentine. See also pitch.

Many pines are also very attractive ornamental trees planted in parks and large gardens. A large number of dwarf cultivars have been selected, suitable for planting in smaller gardens. Some pines are also used for Christmas trees, and pine cones are also widely used for Christmas decorations. Pine trees are also noted for their pleasant smell.

Pine needles serve as food for various Lepidoptera. See List of Lepidoptera which feed on Pines.

Food uses

Main article: Pine nut

Some species have large seeds, called pine nuts, that are harvested and sold for cooking and baking.

The soft, moist, white inner bark (cambium) found clinging to the woody outer bark is edible and very high in vitamins A and C. It can be eaten raw in slices as a snack or dried and ground up into a powder for use as a thickener in stews, soups, and other foods^{[citation needed]}. A tea made by steeping young, green pine needles in boiling water (known as "tallstrunt" in Sweden) is high in vitamins A and C.

Etymology

The modern English name pine derives from Latin Pinus by way of French pin; similar names are used in other Romance languages. In the past (pre-19th century) they were often known as fir, from Old Norse fyrre, by way of Middle English firre. The Old Norse name is still used for pines in some modern north European languages, in Danish, fyr, in Norwegian and Swedish, furu, and Föhre in German, but in modern English, "fir" is now restricted to Fir (Abies) and Douglas-fir (Pseudotsuga).

References

Genera of the Pinaceae family
Pinus · Picea · Cathaya · Larix · Pseudotsuga · Abies · Cedrus · Keteleeria · Pseudolarix · Nothotsuga · Tsuga

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