The rhinoceros (IPA: /raɪˈnɒsərəs/), or
rhino, is any of five surviving species of odd-toed ungulates in the
family Rhinocerotidae. Two species are native to Africa and three to southern Asia. Three of the five species are critically
endangered, and another, the Indian Rhinoceros, is endangered.
The family is characterised by large size (one of the few remaining megafauna surviving
today) with all of the species capable of reaching one ton or more in weight; herbivorous diet; and a thick protective skin, 1.5-5 cm thick, formed from layers of collagen positioned in a lattice structure; relatively small brains for
mammals this size (400-600g); and its horn. The rhino is prized for its horn. The horns of a Rhinoceros are made of keratin, the same type of protein that makes up
hair, but the horn is not itself made of hair as some have believed[1]. Rhinoceros also have acute
hearing and sense of smell, but poor eyesight. Most rhinoceros live to be about 50 years old or more. The collective noun for a group of rhinoceros is "crash".
Both African species and the Sumatran Rhinoceros have two horns, while the Indian
and Javan Rhinoceros have a single horn.
Taxonomy and naming
The word "rhinoceros" (ρινόκερος) is derived from the Greek words rhino, meaning
nose, and kera, meaning horn; hence "horned-nose".
The plural can be rhinoceros, rhinoceri, rhinoceroses, or rhinoceroi. A group of rhinos is called a
crash.
Size comparison of extant rhinoceros species.
The five living species fall into three categories. The two African species, the White
Rhinoceros and the Black Rhinoceros, diverged during the early Pliocene (about 5
million years ago) but the Dicerotini group to which they belong originated in the middle Miocene, about 14 million years ago.
The main difference between black and white rhinos is the shape of their mouths. White rhinos have broad flat lips for grazing
and black rhinos have long pointed lips for eating foliage. The name White Rhinoceros was actually a mistake, or rather a
corruption of the word wijd (wide in Afrikaans) because of their square lips. White Rhinoceros are divided into Northern
and Southern subspecies. There are two living Rhinocerotini species, the endangered Indian
Rhinoceros and the critically endangered Javan Rhinoceros, which diverged from
one another about 10 million years ago. The critically endangered Sumatran
Rhinoceros is the only surviving representative of the most primitive group, the Dicerorhinini, which emerged in the
Miocene (about 20 million years ago).[1] The extinct
Woolly Rhinoceros of northern Europe and Asia was also a member of this tribe.
A subspecific hybrid white rhino (Ceratotherium s. simum × C. s. cottoni) was bred at the Dvůr Králové Zoo (Zoological Garden Dvur Kralove nad Labem) in the Czech Republic in 1977. Interspecific hybridisation of Black and White
Rhinoceros has also been confirmed.[2]
White Rhinoceros
This
White Rhinoceros is actually gray. The origins of the
White in this species
name is uncertain.
-
The White Rhinoceros or Square-lipped Rhinoceros (Ceratotherium simum) is, behind the elephant, probably the most massive remaining land animal in the world, along with the Indian Rhinoceros which is of comparable size and some male hippopotamuses. There are two subspecies of White Rhinos;
as of 2005, South Africa has the most of the first
subspecies, the Southern White Rhino (Ceratotherium simum simum). The population of southern white rhinos is about 11,600,
making them the most abundant subspecies of rhino in the world, and the White Rhino the most abundant species.
The White Rhino has a massive body and large head, a short neck and broad chest. This rhino can exceed 6000 pounds, have a
head-and-body length of 3.35-4.2 m (11-13.9 feet) and a shoulder height of 150-185 cm (60-73 inches). The record-sized White
Rhinoceros was about 3600 kg. On its snout it has two horns. The front horn is larger
that the other horn and averages 89.9 cm (23.6 inches) in length and can reach 150 cm (59 inches). The White Rhinoceros also has
a noticeable hump on the back of its neck which supports its large head. The colour of this animal ranges from yellowish brown to
slate grey. The only hair on them is on the ear fringes and tail bristles. White Rhinos have the distinctive flat broad mouth
which is used for grazing.
Black Rhinoceros
-
The name of the species was chosen to distinguish it from the White Rhinoceros
(Ceratotherium simum). This is confusing, as those two species are not really distinguishable by colour. There are four
subspecies of black rhino: South-central (Diceros bicornis minor), the most numerous, which once ranged from central
Tanzania south through Zambia, Zimbabwe and Mozambique to northern and eastern South Africa;
South-western (Diceros bicornis bicornis) which are better adapted to the arid and semi-arid savannas of Namibia, southern
Angola, western Botswana and western South Africa; East African
(Diceros bicornis michaeli), primarily in Tanzania; and West African(Diceros bicornis longipes) which was
tentatively declared extinct in 2006.[3]
An adult Black Rhinoceros stands 147–160 cm (57.9–63 inches) high at the shoulder and is 3.3-3.6 m (10.8–11.8 feet) in
length.[4] An adult weighs from 800 to 1400 kg
(1,760 to 3,080 lb), exceptionally to 1820 kg (4,000 lb), with the females being smaller than the males. Two horns on the skull are made of keratin with the larger front horn
typically 50 cm long, exceptionally up to 140 cm. Sometimes, a third smaller horn may develop. The Black Rhino is much smaller
than the White Rhino, and has a pointed mouth, which they use to grasp leaves and twigs
when feeding.
Indian Rhinoceros
An Indian rhinoceros and baby at the Nürnberger Zoo.
-
The Indian Rhinoceros or the Great One-horned Rhinoceros (Rhinoceros unicornis) is found in Nepal and in Assam, India. The rhino once
inhabited areas from Pakistan to Burma and may have even roamed
in China. But because of human influence their range has shrunk and now they only exist in small
populations in northeastern India and Nepal. It is confined to
the tall grasslands and forests in the foothills of the
Himalayas.
The Indian Rhinoceros has thick, silver-brown skin which creates huge folds all over its body. Its upper legs and shoulders
are covered in wart-like bumps, and it has very little body hair. Its size is comparable to that of the White Rhino in Africa.
Fully grown males are larger than females in the wild, weighing from 2200–3000 kg (4,800–6,600 lb). Female Indian rhinos weigh
about 1600 kg. The Indian Rhino is from 5.7–6.7 feet tall and can be up to 13 feet long. The record-sized specimen of this rhino
was approximately 3500 kg. The Indian Rhino has a single horn that reaches a length of
between 20 and 101 cm.
Javan Rhinoceros
-
The Javan Rhinoceros (Rhinoceros sondaicus) is one of the rarest and most endangered large mammals anywhere in the world.[5] According to 2002 estimates, only about 60 remain alive, in
Java, Indonesia and Vietnam. Of all the rhino species, the
least is known of the Javan Rhino. These animals prefer dense lowland rain forest, tall grass and reed beds that are plentiful
with large floodplains and mud wallows. Though once widespread throughout Asia, by the 1930's the rhinoceros was nearly hunted to
extinction in India, Burma, Peninsular Malaysia, and Sumatra for the medical powers of its horn
and blood.
Like the closely related larger Indian Rhinoceros, the Javan rhinoceros has only a
single horn. Its hairless, hazy gray skin fall into folds into the shoulder, back, and rump giving it an armored-like appearance.
The Javan rhino's body length reaches up to 3.1-3.2 m (10-10.5 feet), including its head and a height of 1.5–1.7 m tall. Adults
are variously reported to weigh between 900–1,400 kg[6] or
1,360-2,000 kg.[7] Males horns can reach 26 cm in
length while in females they are knobs or no horn at all.[7]
Sumatran Rhinoceros
-
The Sumatran Rhinoceros (Dicerorhinus sumatrensis) is the smallest extant rhinoceros
species, as well as the one with the most fur, which allows it to survive at very high altitudes in Borneo and Sumatra. Due to
habitat loss and poaching, its numbers have declined and it is one of the world's rarest mammals. About 300 Sumatran Rhinos are
believed to remain.
Typically a mature Sumatran rhino stands about 130 cm high at the shoulder, a body length of 240–315 cm and weighs around 700
kg, though the largest individuals have been known to weigh as much as 1,000 kilograms. Like the African species, it has two horns, the largest is the front (25–79 cm) and the smaller being the second which is
usually less than 10 cm long. The males have much larger horns than the females. Hair can range from dense (the most dense hair
in young calves) to scarce. The color of these rhinos are reddish brown. The body is short and has stubby legs. They also have a
prehensile lip.
Evolution
Rhinocerotoids first diverged from other Perissodactyls in at least the Early Eocene. Fossils of Hyrachus eximus found in North America date to this period. This small ancestor resembled a tapir or small
horse, more than a rhino, and had no horn. Three families, sometimes grouped together as the superfamily Rhinocerotoidea, evolved in the Late Eocene: Hyracodontidae, Amynodontidae and
Rhinocerotidae.
Hyracontidae, also known as "running rhinos," showed adaptations for speed, and would have looked more like horses than modern
rhinos. The smallest Hyracontidae were dog-sized; the largest was Indricotherium,
believed to be the largest land-mammal that ever existed. The hornless Indricotherium was almost six meters high, nine meters
long, and weighed as much as 20 tonnes. Like a giraffe, it ate leaves from trees. The Hyracontids spread across Eurasia from the
mid-Eocene to early Miocene.
The Amynodontidae family, also known as "aquatic rhinos," dispersed across North America and Eurasia, from the Late Eocene to
early Oligocene. The Amynodontids were hippopotamus-like in their ecology and appearance, inhabiting rivers and lakes, and
sharing many of the same adaptations to aquatic life as hippos.
The family of all the modern rhinoceroses, the Rhinocerotidae, first appeared in the Late Eocene in Eurasia. The earliest
members of Rhinocerotidae were small and numerous; at least 26 genera lived in Eurasia and North America until a wave of
extinctions in the middle Oligocene wiped out most of the smaller species. Several independent lineages survived, however.
Menoceras, a pig-sized rhinoceros which had two horns side-by-side or the Teloeceras of North America which had short legs and a
barrel chest and lived until about 5 million years ago. The last rhinos in America became extinct during the pliocene.
Modren rhinos are believed to have dispersed from Asia beginning in the Miocene. Two species survived the most recent period
of glaciation and inhabited Europe as recently as 10,000 years ago. The Woolly
Rhinoceros appeared in China around 1 million years ago and first arrived in Europe around 600,000 years ago and again
200,000 years ago, where alongside the Woolly Mammoth, they became numerous but
eventually were hunted to extinction by early humans. Another species of enormous rhino, Elasmotherium, survived the last ice age. Also known as the giant unicorn rhinoceros,
Elasmotherium was two meters tall, five meters long and weighed around five tons, with a single enormous horn, hypsodont
teeth and long legs for running.
Of the extant rhinoceros species, the Sumatran Rhino is the most archaic, first
emerging more than 15 million years ago. The Sumatran Rhino was closely related to the Woolly Rhinoceros, but not to the other
modern species. The Indian Rhino and Javan Rhino are closely related and from a more recent lineage of Asian rhino. The ancestors
of early Indian and Javan rhino emerged 2-4 million years ago.
The lineage of rhinos in Africa is less clear. The black and white rhinoceros remain so closely related that they can still
mate and successfully produce offspring. The black rhinoceros is believed to be the oldest of the species, first emerging between
4 and 10 million years ago, and the white rhinoceros diverging from within the black rhinos between 2 and 5 million years
ago.[8]
- Family Rhinocerotidae
- Subfamily Rhinocerotinae
- Tribe Aceratheriini
- Aceratherium (extinct)
- Acerorhinus (extinct)
- Alicornops (extinct)
- Aphelops (extinct)
- Chilotheridium (extinct)
- Chilotherium (extinct)
- Dromoceratherium (extinct)
- Floridaceras (extinct)
- Hoploaceratherium (extinct)
- Mesaceratherium (extinct)
- Peraceras (extinct)
- Plesiaceratherium (extinct)
- Proaceratherium (extinct)
- Sinorhinus (extinct)
- Subchilotherium (extinct)
- Tribe Teleoceratini
- Aprotodon (extinct)
- Brachydiceratherium (extinct)
- Brachypodella (extinct)
- Brachypotherium (extinct)
- Diaceratherium (extinct)
- Prosantorhinus (extinct)
- Shennongtherium (extinct)
- Teleoceras (extinct)
- Tribe Rhinocerotini
- Tribe Dicerorhinini
- Tribe Ceratotheriini
- Ceratotherium - White
Rhinoceros
- Tribe Dicerotini
- Diceros - Black Rhinoceros
- Paradiceros (extinct)
- Subfamily Elasmotheriinae
- Gulfoceras (extinct)
- Tribe Diceratheriini
- Diceratherium (extinct)
- Subhyracodon (extinct)
- Tribe Elasmotheriini
- Bugtirhinus (extinct)
- Caementodon (extinct)
- Elasmotherium - Giant Unicorn
(extinct)
- Hispanotherium (extinct)
- Huaqingtherium (extinct)
- Iranotherium (extinct)
- Kenyatherium (extinct)
- Menoceras (extinct)
- Ougandatherium (extinct)
- Parelasmotherium (extinct)
- Procoelodonta (extinct)
- Sinotherium (extinct)
Rhinoceros horns
Folk beliefs about rhino horns are a major factor in their decline. (San Diego Zoo)
A rhinoceros horn, believed by some to have aphrodisiac properties.
The most obvious distinguishing characteristic of the rhinos is a large horn above the nose. Rhinoceros horns, unlike those of
other horned mammals, consist of keratin only and lacks a bony core, such as bovine horns.
Rhinoceros horns are used in traditional Asian medicine, and for dagger
handles in Yemen and Oman.
One repeated fallacy is that rhinoceros horn in powdered form is used as an aphrodisiac in Traditional Chinese Medicine. It is, in
fact, prescribed for life-threatening fevers and convulsions and has been clinically shown to have fever-reducing
properties.[9] This misunderstanding has interfered with
discussions with TCM practitioners to reduce its use since the TCM doctors see it as a life-saving medicine of last resort after
cheaper substitutes like water buffalo horn are exhausted.[citation needed] China has signed the
CITES treaty however. To prevent poaching, in certain areas rhinos have been tranquillized and
their horns removed.
Cultural depictions of rhinos
There are a number of legends about rhinoceroses stamping out fire. The story seems to have been
common in Malaysia, India, and Burma. This type of rhinoceros even had a special name in Malay, badak
api, where badak means rhinoceros and api means fire. The animal would come when a fire is lit in the forest
and stamp it out. Whether or not there is any truth to this has not yet been proven, as there has been no documented sighting of
this phenomenon in recent history. This lack of evidence may stem from the fact that rhinoceros sightings overall in
south-east Asia have become very rare, largely due to widespread poaching of the critically endangered animal. This legend is featured prominently in the film
The Gods Must Be Crazy as well as on an episode of The Simpsons.
In the novel James and the Giant Peach by author Roald Dahl, the main character's parents are supposedly eaten by a rhinoceros that had escaped from the
London Zoo. In actuality, Rhinoceros are herbivores.
Albrecht Dürer created a famous woodcut of a
rhinoceros in 1515, without ever seeing the animal depicted. As a result, Dürer's
Rhinoceros is rather inaccurate.
Footnotes
- ^ Rabinowitz, Alan (June 1995) "Helping a Species Go Extinct: The<33
six. Sumatran Rhino in Borneo" Conservation Biology 9(3): pp. 482-488
- ^ Robinson, Terry J.; V. Trifonov, I. Espie,
E.H. Harley (01 2005). "Interspecific hybridization in rhinoceroses: Confirmation of a Black × White rhinoceros
hybrid by karyotype, fluorescence in situ hybridization (FISH) and microsatellite analysis". Conservation Genetics
6 (1): 141-145. doi:10.1007/s10592-004-7750-9.
- ^ "West African black rhino 'is
extinct'", The Times, July 7, 2006. Retrieved on 2007-10-09.
- ^ Dollinger, Peter and Silvia Geser. Black Rhinoceros. World Association of Zoos and
Aquariums. Retrieved on 2007-10-09.
- ^ Derr, Mark.
"[ttp://www.nytimes.com/2006/07/11/science/11rhin.html Racing to Know the Rarest of Rhinos, Before It’s Too Late]",
The New York Times, July 11, 2006. Retrieved on 2007-10-11.
- ^ Species Endangered: Javan Rhinoceros
- ^ a b Rhino Guide: Javan Rhinoceros
- ^ Lacombat,
Frédéric (2005). "The evolution of the rhinoceros", in Fulconis, R.: Save the rhinos: EAZA Rhino Campaign 2005/6. London:
European Association of Zoos and Aquaria,
46-49.
- ^ Chinese Herbal Medicine: Materia Medica, Third Edition, by Dan
Bensky, Steven Clavey, Erich Stoger, and Andrew Gamble. September 2004
References
-
Cerdeño, Esperanza (1995), "Cladistic Analysis
of the Family Rhinocerotidae (Perissodactyla)", Novitates (American Museum of Natural History) (no. 3143), ISSN 0003-0082, <http://digitallibrary.amnh.org/dspace/bitstream/2246/3566/1/N3143.pdf>
- Chapman, January 1999. The Art of Rhinoceros Horn Carving in China. Christies Books, London. ISBN 0-903432-57-9.
-
Emslie, R. and Brooks, M. (1999), African Rhino. Status Survey and Conservation Action Plan.,
IUCN/SSC African Rhino Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK, ISBN 2831705029
-
Foose, Thomas J. and van Strien, Nico (1997), Asian Rhinos – Status Survey and Conservation Action
Plan., IUCN, Gland, Switzerland, and Cambridge, UK, ISBN 2-8317-0336-0
-
Hieronymus, Tobin L. (2006), "Structure of White Rhinoceros (Ceratotherium simum) Horn Investigated by X-ray Computed Tomography and Histology With
Implications for Growth and External Form", Journal of Morphology 267: 1172-1176,
<http://www.oucom.ohiou.edu/dbms-witmer/Downloads/2006_Hieronymus-Witmer-Ridgely_rhino_horn.pdf>
- Laufer, Berthold. 1914. "History of the Rhinoceros." In: Chinese Clay Figures, Part I: Prolegomena on the History of
Defence Armour. Field Museum of Natural History, Chicago, pp. 73-173.
- Parry-Jone, Rob and Amanda Vincent. "Can we
tame wild medicine? To save a rare species, Western conservationists may have to make their peace with traditional Chinese
medicine.", New Scientist, January 3,
1998.
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