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Silurian

 
Dictionary: Si·lu·ri·an   (sĭ-lʊr'ē-ən, sī-) pronunciation
adj.

Of or belonging to the geologic time, system of rocks, or sedimentary deposits of the third period of the Paleozoic Era, characterized by the development of jawed fishes, early invertebrate land animals, and land plants.

n.

The Silurian Period or its system of deposits.

[From Latin Silures, an ancient people of southwest Wales, where the rocks were first identified.]


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Sci-Tech Encyclopedia: Silurian
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The third oldest period of the Paleozoic Era, spanning an interval from about 412 to 438 million years before the present. The Silurian system includes all sedimentary rocks deposited and all igneous and metamorphic rocks formed in the Silurian Period. Both the base and top of the Silurian have been designated by international agreement at the first appearances of certain graptolite species in rock sequences at easily examined and well-studied outcrops. See also Geologic time scale.

The most prominent feature of Silurian paleogeography was the immense Gondwana plate. It included much of present-day South America, Africa, the Middle East, Antarctica, Australia, and the Indian subcontinent. During the Silurian, many plates continued the relative northward motion that had commenced during the mid-Ordovician. Plate positions and plate motions as well as topographic features of the plates controlled depositional environments and lithofacies. These, in turn, significantly influenced organismal development and distributions. Silurian Northern Hemisphere plates, other than a portion of Siberia, are not known north of the Northern Hemisphere tropics. Presumably, nearly all of the Northern Hemisphere north of the tropics was ocean throughout the Silurian. See also Continents, evolution of; Paleogeography.

Absence of plates bearing continental or shallow shelf marine environments north of about 45° north latitude indicates that ocean circulation in most of the Silurian Northern Hemisphere was zonal. Ocean surface currents in the tropics would have been influenced strongly by the prevailing westerlies. The large size of the Gondwana plate and the presence of land over much of it would have led to development of seasonal monsoon conditions. Surface circulation south of 30° south would have hit the western side of Laurentia and flowed generally northward. See also Paleoceanography.

Collision of the Avalonian and Laurentian plates in the latest Ordovician coincides with development of the Southern Hemisphere continental glaciation. Erosion of the land area formed at the Avalon-Laurentian plate collision generated a large volume of coarse to fine-grained siliclastic materials. That part of South America (modern eastern South America) near the South Pole for the early part of the Silurian was the site of as many as four brief glacial episodes. See also Paleoclimatology.

Both nonvascular and vascular plants continued to develop in land environments following their originations in the early mid-Ordovician. Many of these Silurian plants were mosslike and bryophytelike. Psilophytes assigned to the genus Cooksonia were relatively widespread in Late Silurian terrestrial environments. The probable lycopod (club moss) Baragwanathia apparently lived in nearshore settings in modern Australia during the latter part of the Silurian. Silurian land life also included probable arthopods and annelid worms. Fecal pellets of wormlike activity have been found as well as remains of centipede-, millepede-, and spiderlike arthropods. See also Paleobotany; Paleoecology.

Shallow marine environments in the tropics were scenes of rich growths of algae, mat-forming cyanobacteria, spongelike organisms, sponges, brachiopods, bryozoans, corals, crinoids, and ostracodes. Nearshore marine siliclastic strata bear ostracodes, small clams, and snails and trilobites. Certain nearshore strata bear the remains of horseshoe-crab-like eurypterids. See also Algae; Brachiopoda; Bryozoa; Crinoidea; Cyanobacteria; Ostracoda; Trilobita.

Fish are prominent in a number of Silurian nearshore and some offshore marine environments. Jawless armored fish include many species of thelodonts that had bodies covered with minute bony scales, heterostracans, and galeaspids that had relatively heavily armored head shields, and anaspids that possessed body armor consisting of scales and small plates. Jawed fish were relatively rare in the Silurian. They were primarily spiny sharks or acanthodians. As well, there are remains of true sharklike fish and fish with interior bony skeletons (osteichthyans) in Late Silurian rocks. See also Anaspida; Heterostraci; Osteichthyes.


Geography Dictionary: Silurian
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A period of Palaeozoic time stretching approximately from 430 to 395 million years bp.

WordNet: Silurian
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Note: click on a word meaning below to see its connections and related words.

The noun has one meaning:

Meaning #1: from 405 million to 425 million years ago; first air-breathing animals
  Synonym: Silurian period


Wikipedia: Silurian
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Silurian period
443.7 - 416 million years ago
S
SilurianGlobal.jpg
Mean atmospheric O2 content over period duration ca. 14 Vol %[1]
(70 % of modern level)
Mean atmospheric CO2 content over period duration ca. 4500 ppm[2]
(16 times pre-industrial level)
Mean surface temperature over period duration ca. 17 °C [3]
(3 °C above modern level)
Sea level (above present day) Around 180m, with short-term negative excursions[4]

The Silurian is a geologic period and system that extends from the end of the Ordovician period, about 443.7 ± 1.5 Ma (million years ago), to the beginning of the Devonian period, about 416.0 ± 2.8 Ma (ICS, 2004)[5]. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by several million years. The base of the Silurian is set at a major extinction event when 60% of marine species were wiped out. See Ordovician-Silurian extinction events.

Contents

History

The Silurian system was first identified by British geologist Roderick Murchison, who was examining fossil-bearing sedimentary rock strata in south Wales in the early 1830s. He named the sequences for a Celtic tribe of Wales, the Silures, following the convention his friend Adam Sedgwick had established for the Cambrian. In 1835 the two men presented a joint paper, under the title On the Silurian and Cambrian Systems, Exhibiting the Order in which the Older Sedimentary Strata Succeed each other in England and Wales, which was the germ of the modern geological time scale. As it was first identified, the "Silurian" series when traced farther afield quickly came to overlap Sedgwick's "Cambrian" sequence, however, provoking furious disagreements that ended the friendship. Charles Lapworth resolved the conflict by defining a new Ordovician system including the contended beds.

The French geologist Joachim Barrande, building on Murchison's work, used the term Silurian in a more comprehensive sense than was justified by subsequent knowledge. He divided the Silurian rocks of Bohemia into eight stages. His interpretation was questioned in 1854 by Edward Forbes, and the later stages of Barrande, F, G and H, have since been shown to be Devonian. Despite these modifications in the original groupings of the strata, it is recognized that Barrande established Bohemia as a classic ground for the study of the oldest fossils.

Subdivisions

Key events of the Silurian
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-440 —
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-430 —
-425 —
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Key events of the Silurian period.
Axis scale: millions of years ago.

Llandovery

The Llandovery epoch lasted from 443.7 ± 1.5 million years ago to 428.2 ± 2.3 million years ago, and is subdivided into three stages: the Rhuddanian,[8] lasting until 439 million years ago, the Aeronian, lasting to 436 million years ago, and the Telychian.

Wenlock

The Wenlock, which lasted from 428.2 ± 2.3 million years ago to 422.9 ± 2.5 million years ago, is subdivided into the Sheinwoodian (to 426.2 million years ago) and Homerian ages. It is named after the Wenlock Edge in Shropshire, England. During the Wenlock, the oldest known tracheophytes of the genus Cooksonia, appear. The complexity of slightly younger Gondwana plants like Baragwanathia indicates either a much longer history for vascular plants, perhaps extending into the early Silurian or even Ordovician. See Evolutionary history of plants.

Ludlow

The Ludlow, lasting from 422.9 ± 2.5 million years ago to 418.7 ± 2.7 million years ago, comprises the Gorstian stage, lasting until 421.3 million years ago, and the Ludfordian stage.

Přídolí

The Pridoli, lasting from 418.7 ± 2.7 million years ago to 416 ± 2.8 million years ago, is the final and shortest epoch of the Silurian.

Regional stages

In North America a different suite of regional stages is sometimes used:

  • Cayugan (Late Silurian - Ludlow)
  • Lockportian (middle Silurian: late Wenlock)
  • Tonawandan (middle Silurian: early Wenlock)
  • Ontarian (Early Silurian: late Llandovery)
  • Alexandrian (earliest Silurian: early Llandovery)

Silurian paleogeography

Ordovician-Silurian boundary exposed on Hovedøya, Norway, showing the very marked difference between the light gray Ordovician calcareous sandstone and brown Silurian mudstone. The layers have been inverted (overturned) by the Caledonian orogeny.

During the Silurian, Gondwana continued a slow southward drift to high southern latitudes, but there is evidence that the Silurian icecaps were less extensive than those of the late Ordovician glaciation.The southern continents remained united during this period.The melting of icecaps and glaciers contributed to a rise in sea level, recognizable from the fact that Silurian sediments overlie eroded Ordovician sediments, forming an unconformity. Other cratons and continent fragments drifted together near the equator, starting the formation of a second supercontinent known as Euramerica.

Fossilised Late Silurian shallow sea floor, on display in Bristol City Museum, Bristol, England. From the Wenlock epoch, in the Wenlock limestone, Dudley, West Midlands, England.

When the proto-Europe collided with North America, the collision folded coastal sediments that had been accumulating since the Cambrian off the east coast of North America and the west coast of Europe. This event is the Caledonian orogeny, a spate of mountain building that stretched from New York State through conjoined Europe and Greenland to Norway. At the end of the Silurian, sea levels dropped again, leaving telltale basins of evaporites in a basin extending from Michigan to West Virginia, and the new mountain ranges were rapidly eroded. The Teays River, flowing into the shallow mid-continental sea, eroded Ordovician strata, leaving traces in the Silurian strata of northern Ohio and Indiana.

The vast ocean of Panthalassa covered most of the northern hemisphere. Other minor oceans include two phases of the Tethys— the Proto-Tethys and Paleo-Tethys— the Rheic Ocean, a seaway of the Iapetus Ocean (now in between Avalonia and Laurentia), and the newly formed Ural Ocean.

Climate

During this period, the Earth entered a long warm greenhouse phase, and warm shallow seas covered much of the equatorial land masses. Early in the Silurian, glaciers retreated back into the South Pole until they almost disappeared in the middle of Silurian. The period witnessed a relative stabilization of the Earth's general climate, ending the previous pattern of erratic climatic fluctuations. Layers of broken shells (called coquina) provide strong evidence of a climate dominated by violent storms generated then as now by warm sea surfaces. Later in the Silurian, the climate cooled slightly, but in the Silurian-Devonian boundary, the climate became warmer.

Silurian aquatic biota

Artist's impression of Silurian fish

Coral reefs made their first appearance during this time, built by extinct tabulate and rugose corals. The first bony fish, the Osteichthyes, appeared, represented by the Acanthodians covered with bony scales; fishes reached considerable diversity and developed movable jaws, adapted from the supports of the front two or three gill arches. A diverse fauna of Eurypterids (Sea Scorpions) -- some of them several meters in length -- prowled the shallow Silurian seas of North America; many of their fossils have been found in New York State. Leeches also made their appearance during the Silurian Period. Brachiopods, bryozoa, molluscs, hederelloids and trilobites were abundant and diverse.

First terrestrial biota

Cooksonia, earliest vascular plant, middle Silurian

The Silurian was the first period to see macrofossils of extensive terrestrial biota, in the form of moss forests along lakes and streams.

The first fossil records of vascular plants, that is, land plants with tissues that carry food, appeared in the second half of the Silurian period. The earliest known representatives of this group are the Cooksonia (mostly from the northern hemisphere) and Baragwanathia (from Australia). A primitive Silurian land plant with xylem and phloem but no differentiation in root, stem or leaf, was much-branched Psilophyton, reproducing by spores and breathing through stomata on every surface, and probably photosynthesizing in every tissue exposed to light. Rhyniophyta and primitive lycopods were other land plants that first appear during this period. Neither mosses nor the earliest vascular plants had deep roots. Silurian rocks often have a brownish tints, possibly a result of extensive erosion of the early soils.

Some evidence suggests the presence of predatory trigonotarbid arachnoids and myriapods in Late Silurian facies. Predatory invertebrates would indicate that simple food webs were in place that included non-predatory prey animals. Extrapolating back from Early Devonian biota, Andrew Jeram et al. in 1990[9] suggested a food web based on as yet undiscovered detritivores and grazers on microorganisms.[10]

End Silurian extinction

End of Silurian extinction. Click on the picture for more information

At the end of Silurian, a series of minor extinction events, including the Lau event, occurred. They were probably caused by climate change or impact events.[citation needed]

Notes

  1. ^ Image:Sauerstoffgehalt-1000mj.svg
  2. ^ Image:Phanerozoic Carbon Dioxide.png
  3. ^ Image:All palaeotemps.png
  4. ^ Haq, B. U. (2008). "A Chronology of Paleozoic Sea-Level Changes". Science 322: 64–68. doi:10.1126/science.1161648. PMID 18832639. 
  5. ^ Gradstein, Felix M.; Ogg, J. G.; Smith, A. G. (2004). A Geologic Time Scale 2004. Cambridge: Cambridge University Press. ISBN 0521786738. 
  6. ^ Jeppsson, L.; Calner, M. (2007). "The Silurian Mulde Event and a scenario for secundo—secundo events". Earth and Environmental Science Transactions of the Royal Society of Edinburgh 93 (02): 135-154. doi:10.1017/S0263593300000377. 
  7. ^ Munnecke, A.; Samtleben, C.; Bickert, T. (2003). "The Ireviken Event in the lower Silurian of Gotland, Sweden-relation to similar Palaeozoic and Proterozoic events". Palaeogeography, Palaeoclimatology, Palaeoecology 195 (1): 99-124. doi:10.1016/S0031-0182(03)00304-3. 
  8. ^ Named for the Cefn-Rhuddan Farm in the Llandovery area; confusingly, Rhuddlan lies on Silurian strata as well..
  9. ^ Andrew J. Jeram, Paul A. Selden and Dianne Edwards, "Land Animals in the Silurian: Arachnids and Myriapods from Shropshire, England", Science 2 November 1990:658-61.
  10. ^ Anna K. Behrensmeyer, John D. Damuth, et al. Terrestrial Ecosystems Through Time "Paleozoic Terrestrial Ecosystems" (University of Chicago Press), 1992:209.

References

  • Emiliani, Cesare. (1992). Planet Earth : Cosmology, Geology, & the Evolution of Life & the Environment. Cambridge University Press. (Paperback Edition ISBN 0-521-40949-7)
  • Mikulic, DG, DEG Briggs, and J Kluessendorf. 1985. A new exceptionally preserved biota from the Lower Silurian of Wisconsin, USA. Philosophical Transactions of the Royal Society of London, 311B:75-86.
  • Moore, RA, DEG Briggs, SJ Braddy, LI Anderson, DG Mikulic, and J Kluessendorf. 2005. A new synziphosurine (Chelicerata: Xiphosura) from the Late Llandovery (Silurian) Waukesha Lagerstatte, Wisconsin, USA. Journal of Paleontology:79(2), pp. 242-250.
  • Ogg, Jim; June, 2004, Overview of Global Boundary Stratotype Sections and Points (GSSP's) http://www.stratigraphy.org/gssp.htm Accessed April 30, 2006.

External links

Preceded by Proterozoic eon 542 Ma - Phanerozoic eon - Present
542 Ma - Paleozoic era - 251 Ma 251 Ma - Mesozoic era - 65 Ma 65 Ma - Cenozoic era - Present
Cambrian Ordovician Silurian Devonian Carboniferous Permian Triassic Jurassic Cretaceous Paleogene Neogene Quaternary

 
 

 

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