Theodosius Dobzhansky

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Russian–American geneticist (1900–1975)

Dobzhansky, who was born in Nemirov, in Ukraine, graduated in zoology from Kiev University in 1921; he remained there to teach zoology before moving to Leningrad, where he taught genetics. In 1927 he took up a fellowship at Columbia University, New York, where he worked with T. H. Morgan. Morgan was impressed by Dobzhansky's ability and, when the fellowship was completed, offered him a teaching post at the California Institute of Technology. Dobzhansky accepted and became an American citizen in 1937.

Dobzhansky studied the fruit fly (Drosophila) and demonstrated that the genetic variability within populations was far greater than had been imagined. The high frequency of potentially deleterious genes had previously been overlooked because their effects are masked by corresponding dominant genes. Dobzhansky found that such debilitating genes actually conferred an advantage to the organism when present with the normal type of gene, and therefore they tended to be maintained at a high level in the population. Populations with a high genetic load – i.e. many concealed lethal genes – proved to be more versatile in changing environments. This work profoundly influenced the theories on the mathematics of evolution and natural selection with regard to Mendelism.

In addition, Dobzhansky wrote many influential books, including Genetics and the Origin of Species (1937), a milestone in evolutionary genetics.

Theodosius Dobzhansky (1900-1975) synthesized field study and laboratory experimentation in the study of natural selection, laying a foundation for Darwinian evolutionary theory.

Few biologists have made more important contributions to 20th-century evolutionary theory than Theodosius Dobzhansky. His work represents a major part of the synthesis of field study, laboratory experimentation, and classical Mendelian theory that became a powerful foundation for Darwinian theory. Dobzhansky's writings were prolific and influential, comprising over 550 papers and some dozen books. He wrote not only about the technical details of evolution in natural populations, but also about the social and philosophical sides of evolution - including the future evolution of the human species. He lived according to his own dictum, the title of a paper in 1972: "Nothing in biology makes sense except in the light of evolution."

Born on January 25, 1900, in Nemirov, Russia, a small town 200 kilometers southeast of Kiev in the Ukraine, Dobzhansky was the only child of Sophia Voinarsky and Grigory Dobzhansky (sic, the precise transliteration of the Russian name), a high school mathematics teacher. An avid butterfly collector, young Dobzhansky decided to become a biologist at about the age of 12. The family lived on the outskirts of Kiev during the tumultuous years of World War I and the Bolshevik Revolution. During this period Dobzhansky managed to complete his high school and university studies, graduating with a degree in biology from the University of Kiev in 1921.

As an instructor in zoology at the Polytechnic Institute in Kiev (1921-1924), he met Yuri Filipchenko, head of the newly-created Department of Genetics at the University of Leningrad and a strong advocate of T. H. Morgan's work at Columbia University in New York with the small fruit fly, Drosophila melanogaster. Dobzhansky went to work in Filipchenko's lab in 1924, where he began his first studies in genetics. In that same year he married Natalia Sivertzev, a geneticist in her own right working with the famous Russian evolutionist I. I. Schmalhausen in Kiev.

In 1927 Dobzhansky travelled to the United States under the auspices of a fellowship from the International Education Board (Rockefeller Foundation) to work in Morgan's laboratory at Columbia University. Here he began learning the techniques of cytogenetics, particularly the study of chromosome banding structures, that were to be so valuable in his later field studies of evolution in the wild Drosophila population. In September 1928 Dobzhansky moved to the California Institute of Technology, where Morgan had gone to organize and direct the newly-created division of biology. Dobzhansky was named assistant professor of genetics at Caltech in 1929 and professor in 1936. In 1940 he returned to New York as professor of zoology at Columbia University, where he remained for the next 22 years. In 1962 Dobzhansky was appointed professor at the Rockefeller Institute (now Rockefeller University), and in 1971 adjunct professor in the Department of Genetics at the University of California, Davis, a position he held until his death on December 18, 1975.

Dobzhansky's contributions to evolutionary theory relate to five major issues: the amount of variation that exists in natural populations; genetic changes in wild populations due to natural selection; speciation; laboratory studies of fitness under controlled conditions; and human variation and evolution.

Genetic Variation Studies

When Dobzhansky came to the United States in 1927 the predominant view of genetic variation was that established by the work of T. H. Morgan and H. J. Muller in mutant laboratory stocks. Mutations were thought to be relatively rare and other variations in most cases deleterious. Since an organism's overall genetic make-up was the result of natural selection, with deleterious mutations weeded out, wild populations were assumed to harbor few mutations, or variations. As a result, evolution would be - as this was in line with what Darwin had predicted - a relatively slow process.

One of Dobzhansky's major contributions was to show that this view was incorrect. Applying the cytological methods of the Morgan group to the analysis of chromosome structure in wild populations of Drosophila pseudoobscura, Dobzhansky discovered a surprising amount of hidden variability - that is, variations not readily observed in the appearance of individual organisms. Dobzhansky suggested that preservation of extensive variation would allow populations to evolve rapidly as environmental conditions change. Dobzhansky published his findings in Genetics and the Origin of Species in 1937. This book was an important landmark in the evolutionary synthesis: the union of Mendelian genetics and Darwinian theory.

In Morgan's laboratory at Caltech Dobzhansky learned the cytological techniques involved in studying chromosome structure from two of Morgan's most important coworkers, A. H. Sturtevant and C. B. Bridges. In the mid-and late-1930s he had collaborated with Sturtevant on a series of papers using chromosome inversions (where a chromosome segment has been accidentally excised and reinserted in the chromosome upside down) as a way of tracing phylogenetic relationships among species and subspecies of Drosophila. Since inversions are inherited, two separate populations of varieties that showed similarities in chromosome inversion patterns ought to be more closely related than one having fewer similarities.

This work led to a subsequent series of papers of great importance. In the early and mid-1940s, Dobzhansky examined chromosome inversion patterns in populations of Drosophila pseudoobscura from Santa Barbara to central Texas. Each population, he found, had a different frequency for each of several inversion patterns. Moreover, noting that from one season to the next certain inversion patterns increased and others decreased within the same population, Dobzhansky correlated these changes with climatic and other environmental differences associated with changing seasons. He found that one inversion pattern predominated during warmer seasons, while another predominated during colder seasons. Bringing samples from each population back into the laboratory, Dobzhansky showed that he could vary environmental conditions so as to produce the same changes in frequency of inversion patterns that were observed with changing seasons in the field. Dobzhansky concluded that such seasonal fluctuations were the result of natural selection at work, with temperature acting as the selecting agent. These masterful studies provided concrete support for the theory of natural selection, at the same time illustrating the fruitfulness of combining field and laboratory work in the study of evolution.

Speciation and Evolution

As part of the larger question of how speciation occurs, Dobzhansky initiated a number of investigations into the basis of hybrid sterility - that is, the inability of the offspring of many hybrids (especially among animals) to be reproductively fertile. By studying the specifics of sexual, physiological, and behavioral isolating mechanisms in Drosophila pseudoobscura and Drosophila paulistrorum, Dobzhansky showed that varying degrees of reproductive isolation represented speciation in the process. Like many of his other studies, Dobzhansky's work on reproductive isolation was aimed at studying the process of evolution in action.

Although not primarily a human geneticist or paleontologist, Dobzhansky wrote frequently on human evolution, including the biology of race and the future of human evolution. His Mankind Evolving of 1962 was a highly influential work in directing attention to human variation and adaptation. Several subsequent works of a similar nature, such as The Biological Basis of Human Freedom (1956), The Biology of Ultimate Concern (1967), and Genetics of the Evolutionary Process (1970), all reflect Dobzhansky's wide-ranging and philosophical turn of mind.

Although plagued by a form of leukemia in his later years, Dobzhansky remained vigorous and active until the day before his death on December 18, 1975. During his lifetime he was the recipient of many honors and awards. He was a member of the U.S. National Academy of Sciences, the American Academy of Arts and Sciences, the American Philosophical Society, the Royal Society (Great Britain), the Academia Leopoldina (Leipzig), and the Academia Nazionale dei Lincei (Florence). He received the Daniel Giraud Elliot Medal (1946) and the Kimber Genetics Award (1958) from the National Academy of Sciences, the Darwin Medal from the Leopoldina (1959), the A. E. Verrill Award from Yale University (1966), the Gold Medal Award for Distinguished Achievement in Science from the American Museum of Natural History (1969), and the National Medal of Science (1964). In addition, Dobzhansky was awarded honorary degrees by over 20 institutions, including the Universities of São Paulo (Brazil), Münster (Germany), Sydney (Australia), Oxford (England), Padua (Italy), and Chicago, Columbia, Michigan, Syracuse, Berkeley, and Northwestern in the United States.

Further Reading

A detailed biography of Dobzhansky has been prepared by Howard Levene, Lee Ehrman, and Rollin Richmond, "Theodosius Dobzhansky up to now," in Max Hecht and William C. Steere (editors), Essays in Evolution and Genetics in Honor of Theodosius Dobzhansky (1970). A shorter appreciation was written by Francisco Ayala, "'Nothing in biology makes sense except in the light of evolution.' Theodosius Dobzhansky, 1900-1975," in Journal of Heredity (1977). Two important historical essays appear in the reprinted edition of Dobzhansky's "The Genetics of Natural Populations" Series: Richard Lewontin, "Introduction: the scientific work of Th. Dobzhansky," and William B. Provine, "Origins of the 'Genetics of Natural Population' series," in R. C. Lewontin, John A. Moore, William B. Provine, and Bruce Wallace (editors), Dobzhansky's Genetics of Natural Populations I-XLIII (1981). This work contains a complete bibliography.

Theodosius Dobzhansky, ca.1966

Theodosius Grygorovych Dobzhansky, also known as T. G. Dobzhansky, and sometimes Anglicized to Theodore Dobzhansky (Ukrainian — Теодосій Григорович Добжанський; January 24, 1900 - December 18, 1975) was a noted geneticist and evolutionary biologist, and a central figure in the field of evolutionary biology for his work in shaping the unifying modern evolutionary synthesis. Dobzhansky was born in Ukraine (then part of Imperial Russia) and emigrated to the United States in 1927.

Contents 1 Biography 1.1 Early life 1.2 America 1.3 Final illness and the Light of Evolution 1.3.1 Belief in God 2 Bibliography 2.1 Books 2.2 Papers 3 References 4 Further reading 5 Notes 6 External links

Biography

Early life

Dobzhansky was born on January 24, 1900 in Nemyriv, Ukraine (then in the Russian Empire). An only child, his father Grigory Dobzhansky was a mathematics teacher, and his mother was Sophia Voinarsky.^[1] In 1910 the family moved to Kiev, Ukraine. At high school, Dobzhansky collected butterflies and decided to become a biologist.^[2] In 1915, he met Victor Luchnik who convinced him to specialize in beetles instead. Dobzhansky attended the University of Kiev between 1917 and 1921, where he then studied until 1924. He then moved to Leningrad, Russia, to study under Yuri Filipchenko, where a Drosophila melanogaster lab had been established.

On August 8, 1924, Dobzhansky married geneticist Natalia "Natasha" Sivertzeva who was working with I. I. Schmalhausen in Kiev, Ukrainian SSR. The Dobzhanskys had one daughter, Sophie, who later married the American anthropologist Michael D. Coe.

This period was one of great social upheaval in Ukraine and the Russian Empire. The First World War was followed by the Russian Revolution of 1917, and then a civil war that established the Ukrainian Soviet Socialist Republic as a part of the Soviet Union.

America

Dobzhansky emigrated to the United States in 1927 on a scholarship from International Education Board of the Rockefeller Foundation arriving in New York on December 27. He worked with Thomas Hunt Morgan at Columbia University, who had pioneered the use of fruit flies (Drosophila melanogaster) in genetics experiments. He followed Morgan to the California Institute of Technology from 1930 to 1940. Dobzhansky is credited for having taken fruit fly research out of the laboratory and "into the field", having discovered that close regional varieties of flies were more similar to each other genetically than to flies from other regions.

In 1937 he published one of the major works of the modern evolutionary synthesis, the synthesis of evolutionary biology with genetics, entitled Genetics and the Origin of Species, which amongst other things defined evolution as "a change in the frequency of an allele within a gene pool". Dobzhansky's work was instrumental in spreading the idea that it is through mutations in genes that natural selection takes place. Also in 1937, he became a naturalized citizen of the United States. During this time he had a very public falling out with one of his Drosophila collaborators, Alfred Sturtevant, based primarily in professional competition.

Dobzhansky returned to Columbia University from 1940 to 1962. He was one of the signatories of the 1950 UNESCO statement The Race Question. He then moved to the Rockefeller Institute (shortly to become Rockefeller University) until his retirement in 1971. In 1972 he was elected the first president of the BGA (Behavior Genetics Association) ^[3] and was recognised by the society for his role in behavior genetics and the founding of the society by the creation of the Dobzhansky Award (for a lifetime of outstanding scholarship in behavior genetics).

Final illness and the Light of Evolution

Dobzhansky's wife Natasha died of coronary thrombosis on February 22, 1969. Earlier (n June 1, 1968) Theodosius had been diagnosed with lymphocytic leukemia (a chronic form of leukemia), and had been given a few months to a few years to live. He retired in 1971, moving to the University of California, Davis where his student Francisco Jose Ayala had been made assistant professor, and where continued working as an emeritus professor. He published one of his most famous essays "Nothing in Biology Makes Sense Except in the Light of Evolution" at this time.

By 1975, his leukemia had more severe, and on November 11 he travelled to San Jacinto, California for treatment and care. He died (of heart failure) on December 18. He was cremated and his ashes were scattered in the Californian wilderness.

Belief in God

A constant defender of Darwinian evolution and founder of the BGA (Behavior Genetics Association), Dobzhansky, according to Francisco J. Ayala never the less "was a religious man".^[4] Dobzhansky himself spoke of God as creating through evolution, and considered himself a communicant of the Eastern Orthodox Church.

Bibliography

Books

• Sinnott, E.W., Dunn, L.C and Dobzhansky, Th. 1925. Principles of Genetics. McGraw Hill. (4 editions: 1925, 1932, 1939, 1950)
• Dobzhansky, Th. 1937. Genetics and the Origin of Species. Columbia University Press, New York. (2nd ed., 1941; 3rd ed., 1951)
• The Biological Basis of Human Freedom (1954).
• Dunn, L. C., & Dobzhansky, Th. 1946. Heredity, Race, and Society. The New American Library of World Literature, Inc., New York.
• Dobzhansky, Th. 1955. Evolution, Genetics, & Man. Wiley & Sons, New York.
• Dobzhansky, Th. 1962. Mankind Evolving. Yale University Press, New Haven, Connecticut.
• Dobzhansky, Th. 1967. The Biology of Ultimate Concern. New American Library, New York.
• Dobzhansky, Th. 1970. Genetics of the Evolutionary Process. Columbia University Press, New York.
• Dobzhansky, Th. 1973. Genetic Diversity and Human Equality. Basic Books, New York.
• Dobzhansky, Th., F.J. Ayala, G.L. Stebbins & J.W. Valentine. 1977. Evolution. W.H. Freeman, San Francisco.
• Dobzhansky, Th. 1981. Dobzhansky's Genetics of Natural Populations I-XLIII. R.C. Lewontin, J.A. Moore, W.B. Provine & B. Wallace, eds. Columbia University Press, New York. (reprints the 43 papers in this series, all but two of which were authored or co-authored by Dobzhansky)
• Dobzhansky, Th., & Boesiger, E. 1983. Human Culture, A Moment in Evolution. Columbia University Press, New York.

Papers

• Dobzhansky, Th. 1973. "Nothing in Biology Makes Sense Except in the Light of Evolution" The American Biology Teacher 35: (March): 125-129.
• Dobzhansky, T., and O. Pavlovsky. 1957. "An experimental study of interaction between genetic drift and natural selection" Evolution 11: 311-319.

References

• Ayala, Francisco J. (1985). "Theodosius Dobzhansky". Biographical Memoirs of the National Academy of Sciences 55: 163–213. 
• Ford, E. B. (November 1977). "Theodosius Grigorievich Dobzhansky, 25 January 1900 -- 18 December 1975". Biographical Memoirs of Fellows of the Royal Society 23: 58–89. doi:10.1098/rsbm.1977.0004. 

Further reading

• Adams, M. (ed) (1994). The Evolution of Theodosius Dobzhansky : essays on his life and thought in Russia and America. Princeton, N.J.: Princeton University Press. ISBN 0691034796 (acid-free). 

Notes

1. ^ Ford, p59.
2. ^ Ayala, p163.
3. ^ Historical table of BGA Meetingsl
4. ^ Ayala FJ (1976). "Theodosius Dobzhansky: the man and the scientist". Annual Review of Genetics 10: 1–6. doi:10.1146/annurev.ge.10.120176.000245. PMID 797305. 

External links

Wikiquote has a collection of quotations related to: Theodosius Dobzhansky

• Colloquium on Genetics and the Origin of Species with a biography
• Chapter 1 from Genetics and the Origin of Species
• Theodosius Dobzhansky: A Man For All Seasons by Francisco J. Ayala
• The Theodosius Dobzhansky Papers American Philosophical Society

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