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He breaks out of the Templar prison where he was captured in the first game and goes to the Assassins hideout and they also have a animus. He uses the life of Ezio Auditore Da Firenze to learn the ways of the assassins.

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Desmond Miles became an Assassin simply for being born into the Brotherhood of Assassins, though he claimed not to be an Assassin. Desmond was kidnapped by Templar Warren Vidic and his "Fellow Templar" Lucy Stillman who was actually an Assassin in disguise, while in the first Assassins Creed game, Desmond finds out Warren is a Templar, he then escaped with Lucy into the second game.

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