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To understand this saying you need to understand what a lark is.

A lark is a small European songbird which makes its nest in summer in open fields. To attract a mate and to defend its territory, the male lark sings a beautiful, loud, happy sounding, trilling song. It does this by flying high above its nest so, so high that it is difficult to spot the bird even though it is making this lovely noise.

Thus as you hear a lark on nice hot summer days in rolling open country, the sound is associated with summer and happiness.

Thus the expression 'happy as a lark' describes someone 'in their element' perhaps singing away to themselves and obviously very contented with what they are doing.

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Q: What does happier than a lark mean?
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