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That means to get someone to the same level of competency or understanding as others on a team or in a group. It can mean to give someone a briefing or to explain something that a newcomer or latecomer might not already know.
Examples:
"Let me pause in my explanation to bring the latecomers to the meeting up to speed so everyone will know what I am discussing."
"I know you have never used this exact model of machine before, so let me bring you up to speed and brief you on how this machine is different from the ones you have used."
"I don't want to rewind the movie, so I will pause it and bring you up to speed on the plot thus far."
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