It is not an idiom, but polite advice. It was coined in 390AD.
When in Rome, do as the Romans do. This idiom means, when in a new place, try to experience as much of the culture as possible before you leave.
The traditional date for the founding of Rome is 753 BC.
The city of Rome began in 753 BC.
Romans were around since they named their city Rome which was in 753 BC. Last time I checked they were still around today, making the city of Rome and its "Romans" 2,0765 years old.
When in Rome, do as the Romans do.
Rome
The Romans were called Romans because they were from Rome. Roman means people and things from Rome, similarly to English for England.
The Romans were called Romans because they were from Rome. Roman means people and things from Rome, similarly to English for England.
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According to the Romans, Rome was founded by Romulus and Remus.
The date given for the foundation of Rome was 753 BC.
The Romans always called themselves Romans because they were the people from Rome.