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...he discovered the population of mountain yellow-legged frogs (Rana muscosa, Rana sierrae) had declined.

see his published results :

Bradford, D. F., D. M. Graber, and F. Tabatabai. 1994b. Population declines of the native frog, Rana muscosa, in Sequoia and Kings Canyon National Parks, California. Southwestern Naturalist 39:323-327.

Approx a year ago, this frog was added to the list of animals protected under the California Endangered Species Act.

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Q: When David Bradford visited the mountain lakes of Sequoia-Kings Canyon National Parks in the summer of 1988 he discovered that the population of the Rana muscosa had what?
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