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The first air-breathing animals appeared during mid-Palaeozoic era (Era of ancient life) and included invertibrates such as scorpion-like arachnids and wingless insects, both possessing hard exoskeleton. The first reptiles (characterized by hard, dry epidermal scales) diverged from the then-dominant amphibians in the Carboniferous period (345 million years ago) of the Palaeozoic era.

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