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Larvae, to start with, as none of them have wings yet. Also there are two groups of wingless insects; silverfish and fleas (which have lost them along the way, their ancestors had wings). There are quite a few insects that have wings but rarely fly, such as earwigs, most cockroaches, and beetles that have their shields fused so they never fly. Also, in ants the only ones with wings are the queen and males, and the young queen bites off hers when founding her colony to perform better underground. She flies only once, during her mating flight, to get far from her colony of birth to found her own.

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12y ago

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