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Actually, it's the lack of bones. Mammals' jaws have only the dentary (the lower jaw bone) and the squamosal (the upper jaw bone). Non-mammalian amniotes have two more bones in the skull: the articular in the lower jaw and the quadrate in the upper jaw. In them, the articular and the quadrate fuse to form the joint. In mammals, the articular is the malleus of the middle ear, and the quadrate is the incus.

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Q: What bone in the skull is a definitive feature of mammals?
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