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Sequence of Events:a. Light is absorbed by Bchl in the antenna.

b. Energy is transferred to P960 which is a pair of special Bchl b molecules in the reaction center. (P960 means pigment with absorption maximum at 960nm)

c. P960 becomes excited. The excited state, P960*, lasts less than one picosecond (pico = 10-15).

d. P960 loses an electron to one of the other Bchl bmolecules in the reaction center.

e. The electron is rapidly passed on to Bphe. Bphe is a chlorophyll derivative where Mg2+ is replaced by 2H+. The transient existence of the biradical (Bchl)2+....Bphe- is detectable by its spectrum.

f. The electron is transferred from Bphe to firmly bound ubiquinone, UQA, to give the half-reduced form (the anionic semiquinone). [UQA is never fully reduced and never gains protons to become UQH2.]

g. The electron goes from UQA to the loosely bound ubiquinone, UQB. UQB waits for a second electron and also picks up two protons from water so becoming UQH2.

h. UQBH2 moves out of the reaction center, travels through the lipid bilayer, and transfers its electrons to the cytochrome bc1 complex.

i. Cytochrome bc1 transfers electrons to cytochrome c2 which is a soluble periplasmic protein.

j. The electrons are transferred from cytochrome c2 to reaction center protein C and travel successively via its four hemes back to P960+.

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