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This refers to actions taken by a person who, by feel or intuition, with uncontrollable speed, without any assistance from gauges or other devices.

In particular, it refers to a situation where the speaker is overwhelmed and reduced to reacting, rather than planning.
It means to make things up as you go along - to try to do something without any training or instructions. If someone is trying to repair their car without directions, they might say "I'm flying by the seat of my pants here."

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