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It comes from a nursery rhyme, which the protagonist remembers playing with his grandmother:

Wire, briar, linber-lock,

Three geese in a flock.

One flew east, one flew west,

And one flew over the cuckoo's nest.

In this case, cuckoo is also a pejorative term for someone who's crazy or clinically insane. So a cuckoo's nest would be a sanitarium or asylum, where a 'flock' of cuckoos could be found.

Flying over such a nest would be a metaphor for both overseeing the action, and escaping it.

For more info, click the link below, and scroll down to Section II.B: The Meaning of the Title.

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