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The crown eukaryotes are known to have existed in the paleoproterozoic era. Crown eukaryotes are an artificial group of eukaryotic organisms found at the top of molecular phylogenetic trees including both eukaryotes and prokaryotes. They were originally thought to represent a late step of eukaryotic evolution because they include multicellular and macroscopic lifeforms that represent the majority of the biomass of the planet while accounting for less than 1% of the genetic diversity.

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