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It refers to the Service "C" uniform. Dress and Service uniforms in the Marine Corps are designated as "A", "B", "C", or "D" with each version slightly different than the other. Service "A", for example, is green trousers, long-sleeve khaki shirt, tie, and green service coat. Service "C" is green trousers with short-sleeve khaki shirt (no coat). It is called "Chucks" because in the phonetic alphabet the letter "C" is designated as "Charlie". "Charlie" is a nickname for the proper name "Charles" and "Chuck" is a nickname for "Charles", hence the term "Service Charlies" or simply "Chucks".

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