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Mostly just humans, bacteria, viruses, natural conditions. Young cubs are sometimes killed by mature male bears - the mother bear takes great care to protect them from all male bears, sometimes from their own (unknown) father. Sometimes bears are killed by other bears in territorial disputes, but those battles are rarely fatal. A very ill or severely weakened bear might be prey to other large predators, like wolves or cougars, but that's rare, since other predators will have learned to avoid them. Usually if a bear is in any fight with any animal, the opponent will most likely be clawed by the bears razor claws. Also, if the mother is trying to protect the cubs, she will sometimes pick them up in her mouth and run from the male.

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