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That is a good question, the reason for this is that traditionally (in orchestral scores) horns were written for in 2 pairs (when 4 horns were used) often in different keys. So basically you had 1st and 2nd in f and 1st and 2nd (really 3rd and 4th) in Bb, for example (the keys vary depending on the key of the work). This tradition has been carried on into modern music so that usually 1st and 3rd horns have the high parts and 2nd and 4th the low ones. The reason that 1st and 3rd and 2nd and 4th horns are often paired in scores is because their parts are usually more similar.

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