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A song usually begins and starts in the same key, but it doesn't always have to start and end with the same note. However, the last note of a melody can help you guess the key in which that melody was written. For example, if we have a song and it ends with a C, than the key should be C major or minor.

The chord formed on the tonic note -(the tonic being C, in our example) is the most significant chord in a key. To maintain the tonality, our song should finish with the C note (which is the first note of the tonic chord) and start with a note which is contained in the tonic chord (either C, E/Eb, or G). Most of the composers prefer to start with the first note of the chord, so that's why there numerous cases in which the music starts and ends with the same note.

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