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While still attached to the anther, the nucleus of the pollen grain divides, producing two nuclei, one of which becomes the tube nucleus, the other is called the generative nucleus. The generative nucleus divides again, forming two sperm cells.

Once the pollen grain is released from the anther, it is transported (by wind or fauna) to the stigma of the female flower. Once a pollen grain has landed on the stigma, it begins to grow a pollen tube (there are three nuclei in the pollen tube, one from the tube nucleus, and the two sperm). When the pollen tube reaches the ovary, both sperm cells enter the ovule, one fuses with the egg nucleus, the other fuses with polar nuclei that is also inside the ovule, forming the endosperm.

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