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There is no adaptation of cactus plants underwater. The cactus plant is built to withstand light deprivation if it's a jungle cactus. It's built to withstand moisture deprivation if it's a desert cactus. But either way, the cactus can't tolerate excesses in the fulfillment of its moisture, light and heat requirements.

Excess cold and damp soften and blacken cactus tissue. It's an open invitation to fungal problems. No cactus can withstand the large scale softening and blackening of its tissue by being subjected to the constant moisture of underwater living.

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