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Dondake is a slang which originated in the gay area of Shinjuku, and made popular by a gay entertainer named Ikko. Soon high school girls around Harajuku started to use it, and eventually picked up by the mass media. Soon many TV "talentos" started to use it, and even TV doramas. In the recently aired Dorama "Operation Proposal", Johnny's hottie Yamashita Tomohisa (Yamapi) used the word extensively, making it popular even among normal teens.
I'm not clear about the exact origin of Ikahodo, but they have the exact same meaning. The Harajuku teens and possibly the gay population started to use it instead of dondake, and many people have been arguing which is the "correct" or the "cool" term to use. Hence the joke in L*S is that like regular girls, L*S girls also debate about this. Ikahodo is rather new and it's catching up in popularity as the new cool slang to use. The dondake vs ikahodo debate has evolved into the hip dialog of when some one say "Dondake?", you automatically respond with "Ikehodo!".
From: AVPlaya of animesuki.
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