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the sentence isn't grammatically correct and could mean a few things.

1) ¿Por qué no comprende cuando ella habla? means why doesn't he/she/you(formal) understand when she talks?

2) Porque no comprende cuando ella habla means because he/she/you (formal) doesn't understand when she talks.

Did you mean to say "Porque no comprendo cuando ella habla' -- because I don't understand when she talks.

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14y ago

What else can I help you with?