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The earliest tortoise fossils found date to theTriassicperiod.

The remains of a turtle called Odontochelys semitestacea (a half shelled, toothed turtle) have been described from three sets of fossilised remains found in China and dated at around 220 million years old.

As these already show many of the main characteristics of the modern Testudines, it can be reasonably assumed that the ancestors date back further than this.

The probability is that Odontochelys semitestaceaand all modern Testudines evolved from a species of the Anapsids (Clade Anapsida) of the late Carboniferous or early Permian period around 280 - 300 million years ago.

All the Anapsid reptiles, with the exception of Clade Anapsidas and the Procolophonidsbecame extinct during the mass extinction of the Permianperiod.

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