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When a First Sergeant leaves, it'll either be because they've been relieved of duty, or are about to be promoted to Sergeant Major. Who replaces the First Sergeant depends on the availability. It may be a Master Sergeant transferred from a staff position, or it may be a Sergeant First Class who is about to be promoted to E-8. You won't find Master Sergeants in line companies - Master Sergeants are found in staff positions.

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