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The oldest known reptile still alive today is the crocodile.It was an amphibian. There is no major development which i can recall of that allowed them to move on land as they were already capable of doing so. Maybe the origin of terrestrial animals could have led it to find more prey on the land.As according to Darwin's theory of evolution, all animals were first water-borne then came amphibians and then terrestrials and finally avians.
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