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No, watercress is not pruned when it flowers, but just before.
Specifically, the plant in question (Nasturtium officinale) flowers by the time it is 28-31 days old. Cultivators and gardeners generally opt to cut watercress back after 3 weeks. They tend to remove the top 6 inches (15.24 centimeters), save the top 4 inches (10.16 centimeters) of each plant to be used, and trim the remaining plant down to 4 inches in order to encourage multiple harvests.
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