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Is there a silver wolf

Updated: 10/9/2023
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In popular literature, wolf packs are often portrayed as strictly hierarchical social structures with a breeding "alpha" pair which climbs the social ladder through fighting, followed by subordinate "beta" wolves and a low ranking "omega" which bears the brunt of the pack's aggression. This terminology is based heavily on the behaviour of captive wolf packs composed of unrelated animals, which will fight and compete against each other for status. Also, as dispersal is impossible in captive situations, fights become more frequent than in natural settings. In the wild, wolf packs are little more than nuclear families whose basic social unit consists of a mated pair, followed by its offspring.[63] Northern wolf packs tend not to be as compact or unified as those of African wild dogs and spotted hyenas,[64] though they are not as unstable as those of coyotes.[65] Southern wolves are more similar in social behaviour to coyotes and dingoes, living largely alone or in pairs.[66] The average pack consists of 5--11 animals; 1--2 adults, 3--6 juveniles and 1--3 yearlings,[67] though exceptionally large packs consisting of 42 wolves are known. Wolf packs rarely adopt other wolves into their fold, and typically kill them. In the rare cases where strange wolves are adopted, the adoptee is almost invariably a young animal of 1--3 years of age, while killed wolves are mostly fully grown.[68] The adoption of a new member can be a lengthy process, and can consist of weeks of exploratory, non-fatal attacks in order to establish whether or not the newcomer is trustworthy.[69] During times of ungulate abundance (migration, calving etc.), different wolf packs may temporarily join forces.[70] Wolves as young as five months and as old as five years have been recorded to leave their packs to start their own families, though the average age is 11--24 months. Triggers for dispersal include the onset of sexual maturity and competition within the pack for food and breeding.

In areas with low wolf densities, wolves are generally monogamous.[72] Mated pairs usually remain together for life if one of the wolves does not die. Upon the death of one mated wolf, pairs are quickly re-established. Since males often predominate in any given wolf population, unpaired females are a rarity.[73] Polygamy does occur, but primarily in captive situations. Multiple litters are rarely successful, due to infanticide by the pack's females.[74] The age of first breeding in wolves depends largely on environmental factors; when food is abundant, or when wolf populations are heavily managed, wolves can rear pups at younger ages in order to exploit the newly available resources. Captive wolves have been known to breed as soon as they reach 9--10 months, while the youngest recorded breeding wolves in the wild were 2 years old. Females are capable of producing pups every year, with one litter annually being the average. Unlike coyotes, wolves never reach reproductive senescence before they die.[75] Incest rarely occurs, though inbreeding has been reported in Saskatchewan[76] and Isle Royale.[77] Wolf nursing her pups

Estrus typically occurs in late winter, with older, multiparous females entering estrus 2--3 weeks earlier than younger females. Before the rut ensues, wolf packs will temporarily dissolve until the end of the mating season.[73] When receptive, females will avert the base of their tails to one side, exposing the vulva. During mating, the pair is locked into a copulatory tie which may last 5--36 minutes. Because estrus in wolves only lasts a month, the males do not abandon their mates to find other females to inseminate as dogs do. During pregnancy, female wolves will remain in a den located away from the peripheral zone of their territories, where violent encounters with other packs are more likely.[78] Old females usually whelp in the den of their previous litter, while younger females typically den near their birthplace. The gestation period lasts 62--75 days, with pups usually being born in the summer period.[79] The average litter consists of 5--6 pups. Litters of 14--17 occur 1% of the time.[80] Litter sizes tend to increase in areas where prey is abundant.[81] Wolves bear relatively large pups in small litters compared to other canid species.[82] Pups are born blind and deaf, and are covered in short soft grayish-brown fur. They weigh 300--500 grams at birth, and begin to see after 9--12 days. The milk canines erupt after one month. Pups first leave the den after 3 weeks. At 1.5 months of age, they are agile enough to flee from danger. Mother wolves do not leave the den for the first few weeks, relying on the fathers to provide food for them and their young.[82][83] Unlike wolf mothers, the fathers do not regurgitate the pups' food, but carry them pieces from a kill. If the mother dies prior to the pups' weaning period, they are suckled by the pack's other females.[84] Pups begin to eat solid food at the age of 3--4 weeks. Pups have a fast growth rate during their first four months of life: during this period: a pup's weight can increase nearly 30 times.[82][83]

The reproductive behaviour of introduced wolf packs in Yellowstone is unusual, as they often have multiple breeding females who mate with lone male wolves that encroach upon the pack territories during the mating season. These so called "Casanova wolves" are young males that, having failed to procure mates or territories after leaving their natal pack, mate with the daughters of already established breeding pairs from other packs. Unlike males from established packs, Casanova wolves do not form pair bonds with the females they mate with. Because of the great abundance of prey in Yellowstone, female wolves there can bear multiple litters in this fashion.

Wolves use different places for their diurnal rest; places with cover are preferred during cold, damp and windy weather, while wolves in dry, calm and warm weather readily rest in the open. During the autumn-spring period, when wolves are more active, they willingly lie out in the open, whatever their location. Actual dens are usually constructed for pups during the summer period. When building dens, females make use of natural shelters such as fissures in rocks, cliffs overhanging riverbanks and holes thickly covered by vegetation. Sometimes, the den is the appropriated burrow of smaller animals such as foxes, badgers or marmots. An appropriated den is often widened and partly remade. On rare occasions, female wolves will dig burrows themselves, which are usually small and short with 1--3 openings.[86] Wolves do not line their denning places, a likely precaution against parasites.[87] The den is usually constructed not more than 500 metres away from a water source. Resting places, play areas for the pups and food remains are commonly found around wolf dens. The odour of urine and rotting food emanating from the denning area often attracts scavenging birds such as magpies and ravens. As there are few convenient places for burrows, wolf dens are usually occupied by animals of the same family. Though they mostly avoid areas within human sight, wolves have been known to nest near domiciles, paved roads and railways.

Wolves are highly territorial animals, and generally establish territories far larger than they require to survive in order to assure a steady supply of prey. Territory size depends largely on the amount of prey available: in areas with an abundance of prey, the territories of resident wolf packs are smaller. Wolf packs travel constantly in search of prey, covering roughly 9% of their territory per day (average 25 km/d or 15 mi/d). The core of their territory is on average 35 km2 (14 sq mi), in which they spend 50% of their time.[88] Prey density tends to be much higher in the territory's surrounding areas. Despite this higher abundance of prey, wolves tend to avoid hunting in the fringes of their territory unless desperate, due to the possibility of fatal encounters with neighboring packs.[89] The size of their territory may increase when the pack's pups reach the age of 6 months, and thus have the same nutritional requirements as adults. The smallest territory on record was held by a pack of six wolves in northeastern Minnesota, which occupied an estimated 33 km2 (13 sq mi). The largest was held by an Alaskan pack of ten wolves encompassing a 6,272 km2 (2,422 sq mi) area. In some areas, wolves may shift territories during their prey's migration season.[90]

Wolves defend their territories from other packs through a combination of scent marking, direct attacks and howling (see Communication). Scent marking is used for territorial advertisement, and involves urination, defecation and ground scratching. Scent marks are generally left every 240 metres throughout the territory on regular travelways and junctions. Such markers can last for 2--3 weeks,[90] and are typically placed near rocks, boulders, trees or the skeletons of large animals.[91] When scent marking and howling fail to deter strange wolf packs from entering another's territory, violent interactions can ensue.[90] Territorial fights are among the principal causes of wolf mortality: one study on wolf mortality in Minnesota and the Denali National Park and Preserve concluded that 14--65% of wolf deaths were due to predation by other wolves.[92] In fact, 91% of wolf fatalities occur within 3.2 km (2.0 mi) of the borders between neighboring territories.[93] Because the consequences of trespassing can be fatal, such incursions are thought to be largely due to desperation or deliberate aggressiveness.

Although wolf packs do cooperate strategically in bringing down prey, they do not do so as frequently or as effectively as lionesses do; unlike lions, wolves rarely remain with their pack for more than two years, thus they have less time to learn how to hunt cooperatively. Contrary to lion prides, food acquisition per wolf decreases with pack size.[94] Overall, single wolves or mated pairs typically have higher success rates in hunting than do large packs. Single wolves have occasionally been observed to kill large prey such as moose, bison and muskoxen unaided.[95] When hunting, wolves will attempt to conceal themselves as they approach their prey. With ungulate herds, they then either attempt to break up the herd, or isolate one or two animals from it.[96] If the targeted animal stands its ground, the wolves either ignore it, or try to intimidate it into running.[97] When chasing small prey, wolves will attempt to catch up with their prey as soon as possible. With larger animals, the chase is prolonged, in order to wear the selected prey out.[95] Wolves usually give up chases after 1-2 km (0.62-1.3 mi), though one wolf was recorded to chase a deer for 21 km (13 mi).[32] Sometimes, a single wolf will distract the herd with its presence, acting as a decoy, while its pack mates attack from behind.[62] Wolf packs may also set up ambush trails; Indian wolves have been observed to chase gazelle herds through ravines where other wolves lie in wait within holes dug prior to the hunt,[98] while Russian wolves will set up ambushes near water holes, sometimes using the same site repeatedly.[96] Both Russian and North American wolves have been observed to drive prey onto crusted ice, precipices, ravines, slopes and steep banks to slow them down.[99]

Mature wolves usually avoid attacking large prey frontally, instead focusing on the rear and sides of the animal. They kill large prey by biting large chunks of flesh from the soft perineum area, causing massive blood loss. Such bites can cause wounds 10-15 cm in length, with three such bites to the perineum usually being sufficient to bring down a large deer in optimum health.[99] When attacking moose, they occasionally bleed it to death by biting its soft nose.[100] With medium-sized prey such as roe deer or sheep, northern wolves kill by biting the throat, severing nerve tracks and the carotid artery, thus causing the animal to die within a few seconds to a minute.[101] Southern wolves may grab the animal by the neck and stun it by jerking its head downward, hitting its nose on the ground.[10] When prey is vulnerable and abundant, wolves may occasionally surplus kill. Such instances are common in domestic animals, but rare in the wild. In the wild, surplus killing primarily occurs during late winter or spring, when snow is unusually deep (thus impeding the movements of prey)[102] or during the denning period, when wolves require a ready supply of meat when denbound.[103] Medium-sized prey are especially vulnerable to surplus killing, as the swift throat-biting method by which they are killed allows wolves to quickly kill one animal and move on to another.[101] Surplus killing may also occur when adult wolves are teaching their young to hunt.[104]

The breeding pair typically monopolizes food in order to continue producing pups. When food is scarce, this is done at the expense of other family members, especially non-pups.[105][106] This is in marked contrast to the feeding behaviours of dholes and African wild dogs, who give priority to their pups when feeding.[107] The breeding pair typically eats first, though as it is they who usually work the hardest in killing prey, they may rest after a long hunt and allow the rest of the family to eat unmolested. Once the breeding pair has finished eating, the rest of the family will tear off pieces of the carcass and transport them to secluded areas where they can eat in peace. Wolves typically commence feeding by consuming the larger internal organs of their prey, such as the heart, liver, lungs and stomach lining. The kidneys and spleen are eaten once they are exposed, followed by the muscles.

Postural communication in wolves is composed of a variety of facial expressions, tail positions and piloerection.[61] Aggressive or self assertive wolves are characterised by their slow and deliberate movements, high body posture and raised hackles, while submissive ones carry their bodies low, sleeken their fur and lower their ears and tail.[109] When breeding males encounter subordinate family members, they may stare at them, standing erect and still with their tails horizontal to their spine.[110] The pre-caudal scent glands may play a role in expressing aggression, as combative wolves will raise the base of their tails whilst drooping the tip, thus positioning the scent glands at the highest point.[111]

Two forms of submissive behaviour are recognised: passive and active. Passive submission usually occurs as a reaction to the approach of a dominant animal, and consists of the submissive wolf lying partly on its back and allowing the dominant wolf to sniff its anogenital area. Active submission occurs often as a form of greeting, and involves the submissive wolf approaching another in a low posture, and licking the other wolf's face.[112] Howling adult wolf at the UK Wolf Conservation Trust

When wolves are together, they commonly indulge in behaviours such as nose pushing, jaw Wrestling, cheek rubbing and facial licking. The mouthing of each other's muzzles is a friendly gesture, while clamping on the muzzle with bared teeth is a dominance display. Dominant wolves may assert themselves by straddling over a subordinate family member.[113] At a kill, wolves will protect the carcass from afar from other wolves by flattening their ears outwardly, thus indicating that they are covering something belonging to them.

Other vocalisations of wolves are usually divided into three categories: growls, barks and whines.[120] Barking has a fundamental frequency between 320-904 Hz,[24] and is usually emitted by startled wolves. Wolves do not bark as loudly or continuously as dogs do, but will bark a few times and retreat from perceived danger.[120] In captivity, wolves may learn to bark more often if they hear dogs doing so.[121]

Growling has a fundamental frequency of 380-450 Hz,[24] and is usually emitted during food challenges. Pups commonly growl when playing. One variation of the howl is accompanied by a high pitched whine, which precedes a lunging attack.[115] Whining is associated with situations of anxiety, curiosity, inquiry and intimacy such as greeting, feeding pups and playing

Wolf barking at a brown bear in Juraparc, Vallorbe, Vaud, Switzerland

Wolves typically dominate other canid species in areas where they both occur. In North America, incidences of wolves killing coyotes are common, with such incidences being especially common in winter, when coyotes feed on wolf kills. Wolves may attack coyote den sites, digging out and killing the pups. They rarely eat the coyotes they kill. There are no records of coyotes killing wolves, though coyotes may chase wolves if they outnumber them.[133][134] Near identical interactions have been observed in Eurasia between wolves and golden jackals, with the latter's numbers being comparatively small in areas with high wolf densities.[133][135][136] Wolves are the most important predator of raccoon dogs, killing large numbers of them in the spring and summer periods.[137] Wolves also kill red, arctic and corsac foxes, usually in disputes over carcasses. They may eat the foxes they kill.[138][139] In Asia, they may compete with dholes.[140]

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You can get wolves with a silver looking coat, but there's not a silver breed.

http://www.freewebs.com/midnightdreamerpack/SilverWolf.jpg

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