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The composition of their Skeletons.
The Calcarea have single nucleus, single external membrane cells. Their 'Spicule' (Structural Element) is Calcite. They commonly have a massive Calcite exoskeleton
The Glass Sponges cells are mostly syncytia (large structure filled with cytoplasm and many nuclei), and their Spicule is Silica. They never have an exoskeleton.
Finally, the Demosponges have the same calls as the Calcarea, but their Spicule is Silica, like the Glass Sponges. Some species have an exoskeleton, which will be formed of Aragonite if present.
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